Roland Houart
email: roland.houart@skynet.be

Dario Zuccon
email: dario.zuccon@mnhn.fr

Nicolas Puillandre
email: puillandre@mnhn.fr

Cytharomorula Grayi

( Dall

) Mnhn

MD Walters Shoal

Cytharomorula Mk216540

( Grayi

Dall

) Mnhn-Im-2013-66421

Cytharomorula Mk216553

Dw4841 Mnhn

Cytharomorula Mk216548

Dw4789 Mnhn

Cytharomorula Mk216549

Dw4790 Mnhn

Cytharomorula Mk216546

Dw4789 Mnhn

Cytharomorula Mk216544

Dw4789 Mnhn

Cytharomorula Mk216554

Dw4789 Mnhn

Cytharomorula Mk216556

) Mnhn-Im-2013-60932 Karubenthos

Cytharomorula Mk216558

) Mnhn-Im

Mainbaza Cp3130 Mnhn

Cytharomorula Springsteeni

Cp2661 Mnhn

Cytharomorula Vexillum

Ebisco Dw2496 Mnhn

Orania Carnicolor (bozzetti

Dw3608 Mnhn

Orania Mk216552

Carnicolor (bozzetti

Orania Castanea (küster

) Mnhn-Im-2013-66424

Wb10 Mnhn

Orania Mk216555

Castanea (küster

) Mnhn-Im-2013-66431

Ws08 Mnhn

Orania Mk216551

) Mnhn-Im-2013-66432

Ws08 Mnhn

Orania Mk216550

) Mnhn-Im-2013-66434

Wb09 Mnhn

Orania Mk216541

Mnhn-Im-2009-8832 Fischeriana

Miriky

Orania Mixta

Mnhn-Im-2007-18201 Panglao

Orania Pacifica Mnhn

Chinoi N Sp Tenguella

Nhmuk

C Pinguis Houart

C Springsteeni Houart

C Vexillum Kuroda

Maculotriton Dall

M Jacobsoni Emerson & D'attilio

M Rumphiusi Houart

N C Nc Nc

Spinidrupa Habe

Kosuge

Murex Euracanthus A Adams

Description of new genera and new species of Ergalataxinae (Gastropoda: Muricidae)

Keywords: New taxa, Gastropoda, Muricidae, Ergalataxinae, Indo-Pacific, South Atlantic

The recent genetic analysis of the muricid subfamily Ergalataxinae has led to a better understanding of this subfamily, but some species were left without appropriate generic assignments and the classification of others required revision. This knowledge gap is partially filled herein, with new combinations and the description of three new genera. The examination of new material, along with a careful re-examination of and comparison to existing material, resulted also in the identification of nine new species. These new genera and new species are described herein, lectotypes are designated and new combinations are given. The geographical range of all the new species is provided on maps. All new species are compared with related or similar species. The radula of Morula palmeri Powell, 1967 is illustrated for the first time. New genera:

INTRODUCTION

Four species were originally assigned to the subfamily Ergalataxinae (Gastropoda, Muricidae): Bedevina birileffi (Lischke, 1871), Ergalatax contracta [START_REF] Reeve | Conchologia iconica , or illustration of the shells of molluscous animals[END_REF], E. tokugawai Kuroda & Habe, 1971 and Cytharomorula vexillum Kuroda, 1953. The subfamily diagnosis was based on morphological characters of the shell, radula and egg capsule [START_REF] Kuroda | The Sea Shells of Sagami Bay[END_REF]). Later on, many genera and species were added to the subfamily, counting now 20 genera and 146 species recognized as valid (WORMS). Recent molecular genetic analyses confirmed the validity of this subfamily within the Muricidae [START_REF] Barco | A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods[END_REF][START_REF] Claremont | A molecular phylogeny of the Rapaninae and Ergalataxinae (neogastropoda: Muricidae)[END_REF][START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF]. [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] analysed 52 ergalataxine species in 18 genera, representing 36% of the currently accepted species and 90% of the genera, and showed that many of the currently accepted genera were polyphyletic, thus requiring further analysis. In addition, some of the species included by [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] were unidentified, and potentially represented new species. To address some of these issues, we herein discuss and clarify the taxonomy of the genera Muricodrupa Iredale, 1918, Cytharomorula Kuroda, 1953, Orania Pallary, 1900, Pascula Dall, 1908, and Tenguella Arakawa, 1965. In addition, three new genera are described, together with several new species in Cytharomorula, Orania, Pascula, Tenguella and Claremontiella n. gen. Our conclusions are based on the results from Claremont et al 2013, completed by the examination of additional material, including newly sequenced samples.

Material and methods

Material

The material studied here primarily includes specimens collected on various cruises conducted by the MNHN/IRD in the Indo-West Pacific. Other specimens were from the collections of the Natural History Museum, United Kingdom; the KwaZulu-Natal Museum, South Africa; the Houston Museum of Natural History, Houston, Texas, U.S.A; and the personal collection of the first author.

Specimens from the following expeditions of the MNHN/IRD were examined: BENTHEDI: Mayotte, northern Madagascar, Mozambique Channel, 1977;MD32: Reunion Island, 1982;BIOCAL: New Caledonia, 1985;CHALCAL 2: New Caledonia, 1986;SMIB 3: New Caledonia, 1987;SMIB 4: New Caledonia, 1989;SMIB 5: New Caledonia, 1989;BERYX 11: New Caledonia, 1992;MUSORSTOM 7: SW Pacific, Wallis & Futuna, 1992;SMIB 8: New Caledonia, 1993;BATHUS 2: New Caledonia, 1993;MUSORSTOM 8: Vanuatu, 1994; MUSORSTOM 9: Marquesas, 1997;LIFOU 2000: New Caledonia, 2000;BORDAU 2: Tonga, 2000;NORFOLK 1: New Caledonia, 2001;NORFOLK 2: New Caledonia, 2003;PANGLAO 2004 and2005: Philippines;EBISCO: New Caledonia, 2005;SANTO 2006: Vanuatu, 2006;AURORA 2007: Philippines;TERRASSES: New Caledonia, 2008;MAINBAZA: Mozambique, 2009;MIRIKY: Madagascar, 2009; ATIMO VATAE: Madagascar, 2010; EXBODI: New Caledonia, 2011; PAPUA NIUGINI: Papua New Guinea, 2012; PAKAIHI I TE MOANA: Marquesas, 2012;KANACONO: New Caledonia, 2016;KANADEEP: New Caledonia, 2017;MD208: Walters Shoal, south of Madagascar, 2017 andBIOMAGLO: Mayotte, Glorieuses andComoros islands, 2017. The material used for the phylogenetic tree is listed in Table 1 Morphological analyses

The characters used to describe shell morphology address the general aspect of the shell, its shape, size, and colour, the shape of the spire including the number and features of the protoconch and teleoconch whorls, details of the suture and of the subsutural ramp, details of axial and spiral sculpture, the aperture, the siphonal canal, and when available, the characters of the operculum and radula.

The method used to determine diameter and height, and to count the number of protoconch whorls, follows [START_REF] Bouchet | New Caledonia: the major center of biodiversity for volutomitrid mollusks (Mollusca: Neogastropoda: Volutomitridae)[END_REF] as shown in Fig. 1. The morphology of the radula is described starting from the rachidian tooth, followed by the lateral teeth (Fig. 2). Unless otherwise indicated, species descriptions are based on the holotype and the paratypes. The bathymetric ranges given herein are the inner values of the recorded depths: the deepest minimum and the shallowest maximum of each recorded depth range.

Molecular analyses

The protocols used for DNA extraction and the PCR and sequencing of the Barcode fragment of the Cytochrome oxidase I (COI) gene are detailed in [START_REF] Puillandre | A quest for the lost types of Lophiotoma (Gastropoda: Conoidea: Turridae): integrative taxonomy in a nomenclatural mess[END_REF]. PCR products were purified and sequenced by the Eurofins sequencing facility. Sequences were deposited in BOLD (Barcode of Life Datasystem) and GenBank (Table 1). Newly obtained sequences were combined in a single dataset together with the sequences from [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] that corresponds to the same genera, i.e. Cytharomorula, Orania and Tenguella and two outgroups (Concholepas concholepas and Thais nodosa) (Table 1).

All the sequences were aligned manually (no indel was detected), and the dataset was analysed using a Bayesian approach as implemented in Mr.Bayes v. 3.2 [START_REF] Huelsenbeck | MrBayes: Bayesian inference of phylogeny[END_REF], with two runs consisting of four Markov chains of 10,000,000 generations each, with 8 chains, 5 swaps, and a sampling frequency of one tree every 2,000 generations. Each codon position of the COI gene was treated as an unlinked partition, each following a general time reversible (GTR) model, with a gamma-distributed rate variation across sites approximated in four discrete categories and a proportion of invariable sites. Convergence of each analysis was evaluated using Tracer v. 1.6 [START_REF] Rambaut | Tracer v1.6[END_REF] to check that all ESS values were greater than 200. The trees were then calculated after omitting the first 25% trees as burnin. Statistical support was evaluated as Bayesian posterior probability (PP). Convex part of teleoconch whorl and siphonal canal ab: abapical (or abapertural); abis: abapical infrasutural secondary cord on subsutural ramp; ABP: abapertural primary cord on the siphonal canal; ad: adapical (or adapertural); adis: adapical infrasutural secondary cord on subsutural ramp; ADP: adapertural primary cord on the siphonal canal; ads: adapertural secondary cord on the siphonal canal; IP: infrasutural primary cord on subsutural ramp; MP: median primary cord on the siphonal canal; ms: median secondary cord on the siphonal canal; P: primary cord; P1: shoulder cord; P2-P6: primary cords of the convex part of the teleoconch whorl; s: secondary cord; s1-s6: secondary cords of the convex part of the teleoconch whorl (example: s1 = secondary cord between P1 and P2; s2 = secondary cord between P2 and P3, etc.); SP: subsutural cord.

Results and Discussion

Molecular analyses

In the phylogenetic tree (Fig. 3), the genera Orania and Tenguella are recovered monophyletic. Cytharomorula is recovered paraphyletic, contrary to the results obtained by [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], but this is probably due to the fact that we used only one gene. Each included species, and in particular, the newly described species, Cytharomorula absidata n. sp., Cytharomorula elegantula n. sp., Tenguella chinoi n. sp. and Tenguella ericius n. sp., corresponds to an independent lineage, and when several specimens per species are included, they are grouped in a single supported clade (Posterior Probabilities > 0.95).

Taxonomic implementation

Given the results obtained in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], the phylogenetic tree presented here (Fig. 3) and the morphological analyses of additional material (see below), we propose to revise several genera of Ergalataxinae and to describe new genera and new species. The different taxa are discussed below in the order they appear in the fig. 1 of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], following their clade naming (A, B, C, V, W, X, Y and Z); it should be noted that the clades for which we do not propose any taxonomic change or novelty are not discussed.

Purpura nodulosa C.B. Adams, 1845 has previously been assigned to Morula Schumacher, 1817 or Trachypollia Woodring, 1928 by a few authors: to Morula by [START_REF] Rios | Coastal Brazilian seashells[END_REF][START_REF] Rios | Seashells of Brazil[END_REF], [START_REF] Abbott | American Seashells[END_REF], [START_REF] Bernard | Coquillages du Gabon[END_REF] and [START_REF] Houart | Les Muricidae d'Afrique Occidentale -II. Ocenebrinae, Ergalataxinae, Tripterotyphinae, Typhinae, Trophoninae & Rapaninae[END_REF]; and to Trachypollia by [START_REF] Radwin | Murex shells of the world. An illustrated guide to the Muricidae[END_REF], [START_REF] De | Marine gastropods from Curaçao, Aruba and Bonaire[END_REF], [START_REF] Leal | Marine Prosobranch Gastropods from Oceanic Islands off Brazil: species composition and biogeography[END_REF] and Diaz [START_REF] Diaz Merlano | Moluscos del Caribe Colombiano Un catâlogo ilustrado[END_REF]. A sister relationship between "Morula" nodulosa and all other species in major clade A was well supported in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], who suggested to create a new genus for the western and eastern Atlantic representatives of this clade.

"Morula" nodulosa was previously included in Morula because its shell morphology was similar to that of species of Tenguella, previously considered a synonym of Morula. However, molecular genetic studies have shown this species to be separated from both Tenguella and Morula, as well as from Trachypollia. A new genus is here described to include "Morula" nodulosa and two other species.

Muricodrupa has long been considered to contain three species: the type species M. fenestrata [START_REF] Blainville | Disposition méthodique des espèces récentes et fossiles des genres Pourpre, Ricinule, Licorne et Concholepas de M. de Lamarck, et description des espèces nouvelles ou peu connues, faisant partie de la collection du Muséum d'Histoire Naturelle de Paris[END_REF] (Fig. 29A-B), M. fiscellum [START_REF] Gmelin | Caroli a Linné Systema Naturae per Regna Tria Naturae, Editio decima tertia, aucta, reformata[END_REF] (Fig. 28F-H) and M. jacobsini Emerson & D'Attilio, 1985 (Fig. 28I). However, based on [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], M. fiscellum should be excluded from Muricodrupa, probably together with M. jacobsini, a different although morphologically related species. Muricodrupa fenestrata would then remain as the sole representative of this genus. However, [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] noted that "Morula" anaxares [START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF] formed a marginally significant clade with M. fenestrata, which was not contradicted in any other analyses. Indeed, the shell morphology of "Morula" anaxares is very similar to M. fenestrata (Fig. 29C-D). As it is clear that "Morula" anaxares is excluded from Morula and Tenguella, we here include this species in Muricodrupa.

In the molecular analysis of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], M. fiscellum formed a well-supported clade with "Morula" rumphiusi Houart; 1996, which was clearly excluded from Morula Schumacher, 1817. Both species, together with M. jacobsini, are morphologically distinct from the other species included in the clade B [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF]), and we here assigned them to a new genus (see Table 2).

Eleven species typically assigned to Morula formed a monophyletic group within major clade B (subclade V) of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF]. Since this subclade included M. uva, the type species of Morula, Claremont et al (2013: 27) proposed that this generic name should be restricted to this subclade. This subclade also contained M. biconica, the type species of Habromorula Houart, 1995, but the Habromorula species analysed by [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] and assigned to this group [as a subgenus of Morula: M. (H.) biconica, M. (H.) coronata, M. (H.) japonica, M. (H.) spinosa and M. (H.) striata] did not form a clade in any analysis. However, the authors also noted that the monophyly of this morphologically distinctive group was not strongly contradicted, so further sampling and analysis could yet support Habromorula as a monophyletic subgenus of Morula.

Three species currently assigned to Orania, O. gaskelli (Melvill, 1891) (Fig. 21L), O. serotina (A. Adams, 1853) (Fig. 21M-N) and O. bimucronata [START_REF] Reeve | Conchologia iconica , or illustration of the shells of molluscous animals[END_REF] (Fig. 21Q) formed the subclade W in the molecular phylogeny of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], clustering with Lataxiena fimbriata (Hinds, 1843) (Fig. 22J) and Usilla avenacea (Lesson, 1842) (Fig. 21O-P), and not with the other species of Orania (clade Z, see below). Based on shell morphology, it is certainly possible to group O. gaskelli, O. serotina and possibly U. avenacea in the same genus. However, it would be difficult to justify the inclusion of O. bimucronata and particularly L. fimbriata in that morphological group, and additional studies are required before grouping them in the same genus. We suggest continuing to retain these species in the genera to which they are currently assigned: Orania (O. gaskelli, O. serotina and O. bimucronata), Lataxiena (L. fimbriata) and Usilla (U. avenacea) (see Table 2).

The type species of Oppomorus Iredale, 1937, O. nodulifera (Menke, 1829), formed a well-supported clade with O. funiculata [START_REF] Reeve | Conchologia iconica , or illustration of the shells of molluscous animals[END_REF] and O. purpureocincta (Preston, 1909), and [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] therefore proposed that Oppomorus should be accorded full generic rank and not considered a subgenus of Morula, as had been previously proposed by Houart (2004). Three species formerly assigned to Morula [M. nodulosa (C.B. [START_REF] Adams | Speciorum novarum conchyliorum Jamaica repertorum synopsis[END_REF], M. parva (Reeve, 1845) and M. rumphiusi Houart, 1996] were excluded from the genus.

In major clade C of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], "Morula" parva [START_REF] Reeve | Conchologia iconica , or illustration of the shells of molluscous animals[END_REF] was sister to all other members of subclade Z, all morphologically clearly distinct from "Morula" parva, indicating that a new genus was also required for this species.

The genus Pascula was also recovered as polyphyletic by [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF], and is included in their Clade C, subclade Z, together with Cytharomorula and Orania. The type species of Pascula, P. citrica (Dall, 1908) (Fig. 22K-L) was not analysed in that study. However, the study did include a related species, P. darrosensis (E.A. Smith, 1884), with a similar radula morphology . Pending further genetic research all species previously included in Pascula are here retained in this genus (see Table 2).

Based on the results of the present analysis, and also because the shell morphology of the type species of Orania [(Orania fusulus (Brocchi, 1814)], as well as that of other Orania species, is similar to the shell morphology of Cytharomorula, the latter could be considered a possible junior synonym of Orania, pending future research. Indeed, Claremont et al. (2013: subclade Z) considered the validity of Cytharomorula uncertain, as this genus was not monophyletic in any analysis. However, this nonmonophyly was due to the presence of Orania ornamentata Houart, 1995 (Fig. 13D-E) in the Cytharomorula clade. The shell morphology of O. ornamentata is strongly similar to that of the type species of Cytharomorula (C. vexillum Kuroda, 1953) (Fig. 13A-C), and we thus re-assign O. ornamentata in Cytharomorula. This renders the Cytharomorula of [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] subclade Z) monophyletic. Unfortunately, as long as Orania fusulus (Brocchi, 1814) (Fig. 21A-C), the type species of Orania, has not been analysed genetically, it will remain unclear whether Cytharomorula should be considered a junior synonym of Orania or not. Here, we still consider that Orania and Cytharomorula are two distinct genera.

Four ergalataxine genera were not included in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] because of a lack of appropriate material, and the type species were not analysed for two additional genera (Table 2). The inclusion of these four genera in the Ergalataxinae, and the composition of Orania and Pascula, therefore remains doubtful. Their classification is primarily based on morphological shell and/or radula characters.

SYSTEMATIC ACCOUNT

Family MURICIDAE Rafinesque, 1815 Subfamily ERGALATAXINAE [START_REF] Kuroda | The Sea Shells of Sagami Bay[END_REF] Genus Cytharomorula Kuroda, 1953 Type species by monotypy: Cytharomorula vexillum Kuroda, 1953, Japan (Fig. 13A-C) Kuroda, 1953 (Pacific Ocean). Only one of these species occurs in the Atlantic, all others are found in the Indo-West Pacific region. A revision of a few species (Cytharomorula ambonensis, C. benedicta, C. dollfusi, C. lefevreiana and C. paucimaculata) was recently published (Houart, 2013a). Five additional new species from the Indo-West Pacific are described below, and a new combination is given for C. ornamentata [START_REF] Houart | The Ergalataxinae (Gastropoda, Muricidae) from the New Caledonia region with some comments on the subfamily and the description of thirteen new species from the Indo-West Pacific[END_REF], which gives us a total of 16 species for this genus.

In several species the primary spiral cord P4 is generally broader and larger, with broader, more conspicuous knobs at the intersection with the axial ribs. Cytharomorula lefevreiana (Fig. 15F-G), another species occurring off Mauritius and Reunion that was collected during the MD32 cruise (Houart, 2013a), differs in many aspects, including having a comparatively smaller and narrower shell, from 7 to 11 mm in length, more strongly shouldered whorls with very obvious P1-P3 spiral cords, and a relatively smaller aperture.

Cytharomorula absidata

Etymology. Arta (L): meaning narrow, named for its narrow shell.

CYTHAROMORULA GRAYI GROUP

A very careful comparative study of shell and genetic characters, including additional newly available material, indicates that specimens identified or doubtfully identified as C. grayi [START_REF] Dall | Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877-78) and in the Caribbean Sea (1879-80), by the U.S. Coast Survey Steamer "Blake"... 29. Report on the Mollusca. 2, Gastropoda & Scaphopoda[END_REF] was "removed" from its box in the MCZ, and the specimen presently in the box labelled C. grayi is not C. grayi but another species, not even a muricid [START_REF] Tröndlé | Les Muricidae de Polynésie Française[END_REF][START_REF] Vokes | Cenozoic Muricidae of the western Atlantic region. Part XI -The subfamily Ergalataxinae[END_REF], but rather a buccinid. Nevertheless, the original figure in Dall (1889: pl. 32, fig. 12a) is excellent and its identity is undoubtable. The specimen illustrated by Dall was designated as the lectotype by Vokes (1996: 29); [START_REF] Watson | On the marine Mollusca of Madeira; with descriptions of thirty-five new species, and an index-list of all the known seadwelling species of that island[END_REF]: lectotype NHMUK 1911.7.17.2 (Fig. 15N), designated by Houart & Abreu (1994: 122) (see remarks).

Cantharus (Tritonidea) laevis E.A.
Distribution. Cytharomorula grayi occurs from the Lesser Antilles in 100-450 m [START_REF] Garrigues | Muricidae récoltés au cours de la campagne KARUBENTHOS 2 du MNHN dans les eaux profondes de Guadeloupe (Antilles Françaises) et description de trois nouvelles espèces des genres Pagodula et Pygmaepterys (Mollusca, Gastropoda)[END_REF] to southern Brazil in the western Atlantic, and in the eastern Atlantic in the Canary Islands, Madeira and the banks off Portugal, but not on the mainland European continental shelf (SEAMOUNT 1 cruise material in MNHN). The species also occurs off Saint Helena and extends its range to South Africa, Walters Shoal, Mozambique, Reunion, Mayotte, Comoros and Glorieuses Islands, Indian Ocean, there living in 128-275 m (Fig. 14).

Remarks. Vokes (1996: 29) mentioned and illustrated the originally figured syntype of Cantharus laevis as the holotype. Smith (1890) never designated any shell as the holotype. The type material in the NHMUK consists of 5 specimens, mentioned as syntypes by Houart (1997: 56). [START_REF] Vokes | Cenozoic Muricidae of the western Atlantic region. Part XI -The subfamily Ergalataxinae[END_REF] only mentions what she then called the "holotype", but having examined the material when visiting the NHMUK she certainly was aware of the existence of the other 4 specimens. The mention of Vokes is here considered as an indirect lectotype designation by inference as holotype, following ICZN Art. 74.5: "In a lectotype designation made before 2000, either the term "lectotype", or an exact translation or equivalent expression (e.g. "the type"), must have been used or the author must have unambiguously selected a particular syntype to act as the unique name-bearing type of the taxon. When the original work reveals that the taxon had been based on more than one specimen, a subsequent use of the term "holotype" does not constitute a valid lectotype designation unless the author, when wrongly using that term, explicitly indicated that he or she was selecting from the type series that particular specimen to serve as the namebearing type". Houart & Abreu (1994: 122, fig. 5) illustrated the syntype and only existing specimen of Trophon lowei as the holotype. This is here also considered as a valid lectotype designation following ICZN Art. 74.6: "When it has been accepted that a nominal speciesgroup taxon was based on a single specimen and the original description neither implies nor requires that there were syntypes, and if it is considered subsequently that the original description was based on more than one specimen, the first author to have published before 2000 the assumption that the species-group taxon was based upon a single type specimen is deemed to have designated that specimen as the lectotype".

Discussion. Material consisting of numerous live and dead collected specimens from the southwestern Indian Ocean was examined and initially considered a separate species. Adult specimens are bigger than the typical Atlantic Cytharomorula, wider and more shouldered, have a slightly wider aperture and a narrower siphonal canal (Figs 11A-B; 16A-G). Nevertheless, genetic analysis performed on specimens from Guadeloupe (KARUBENTHOS 2), Walters Shoal (MD 208) and other specimens collected off Mayotte, Comoros and Glorieuses islands confirm the conspecifity of the populations of the Atlantic Ocean and the Indian Ocean. The type material of Trophon lowei (Madeira) (Fig. 15N) and Cantharus laevis (Saint Helena) (Fig. 15O-P) seems to be composed of intermediate forms between the populations of the western Atlantic and the Canary Islands (Fig. 15H-L) and those of the Indian Ocean.

Cytharomorula elegantula n. sp.

Figs 6B-C; 10C; 14; 16H-N Cytharomorula grayi -Houart, 1995: 254, figs 9-10, 67-68;[START_REF] Houart | Muricidae (Mollusca: Gastropoda) from Fiji and Tonga[END_REF]: 459 [(not C. grayi (Dall, 1889)]. Distribution. New Caledonia, Vanuatu, Futuna and Tonga, living at 100-505 m, but most living specimens were collected between 230 and 265 m (Fig. 14). Cytharomorula springsteeni (Fig. 13M-O) also differs from C. manusuduirauti n. sp. in many ways. The shell of C. springsteeni is narrower and more elongate, with a larger, higher aperture and a higher spire. The spiral sculpture also differs in being more irregular with higher, weakly broader primary and secondary cords compared to the narrower tertiary cords (Fig. 10F), rather than the similar sized, more numerous and crowded cords in C. manusuduirauti n. sp.

Description

Etymology. Named for the late Emmanuel (Manu) Guillot de Suduiraut, friend and well-known shell enthusiast who sent a specimen (now paratype) of this new species to one of the authors (RH) several years ago.

Genus Orania Pallary, 1900 Type species by original designation: Pseudomurex spadae Libassi, 1859 (= Murex fusulus Brocchi, 1814), Mediterranean (Fig. 21A-C). 2). The radula of Orania fusulus (Fig. 6G), the type species, is typical for the Orania type described in Houart (1995: 246, fig. 3), as it has broader cusps and denticles, small, short marginal cusps and a short marginal cusp. See also the radula of Orania carnicolor (Fig. 6E-F).

Diagnosis

Orania pseudopacifica

Distribution.

French Polynesia, Marquesas Archipelago, 7°48'S -10°29'S, 138°36'W -140°44'W, living at 50-117m (Fig. 19).

Description. Shell medium sized for the genus, up to 16.8 mm in length at maturity (paratype MNHN). Length/width ratio 1.7-1.8. Biconical, broadly-ovate, heavy, weakly nodose, finely squamous. Subsutural ramp broad, strongly sloping, weakly concave. White, light tan or light peach. Aperture white, occasionally tinged with light pink at edge of columellar lip. Spire high with 3.5-4 protoconch whorls and teleoconch of up to 5 broad, weakly shouldered whorls. Suture of whorls adpressed. Protoconch small, conical, acute. Whorls smooth, glossy, with a very narrow, single keel abapically. Height 900 µm, width 750-800 µm. Terminal lip of sinusigera type.

Axial sculpture of teleoconch whorls consisting of low, broad, rounded ribs. First and second teleoconch whorls with 7-9 ribs, third with 8 or 9, fourth with 7 or 8, last whorl with 6 or 7 ribs. Spiral sculpture of high, rounded, narrow, finely squamous, primary, secondary and tertiary cords and few additional narrow threads. First whorl with SP, IP, P1, second whorl with SP, IP, starting abis, P1, s1, third with SP, starting adis, IP, abis, P1, t, s1, fourth with SP, adis, IP, abis, P1, t, s1, last teleoconch whorl usually with SP, adis, IP, abis, P1, t, s1, t, P2, s2, t, P3, s3, P4, s4, t, P5, s5, ADP, MP and one or two additional threads. SP sometimes split, broad, strongly squamous, occasionally with high, projecting scales. Aperture moderately large, narrow, ovate. Columellar lip narrow with 2-4 strong knobs on whole length, deeply extended within aperture (Fig. 11D). Rim partially erect, adherent at small portion adapically. Occasionally with weak, low parietal tooth at adapical extremity. Anal notch deep, broad, with expanded lip. Outer lip erect, crenulated, with 6 strong, elongate denticles within, decreasing in strength abapically, consisting of ID, D1-D5. Siphonal canal short, broad, dorsally recurved, broadly open. Radula of three-dimensional type consisting of a rachidian with a narrow, long central cusp, a short lateral denticle and a fairly broad, moderately long lateral cusp on each side, few, obvious, marginal folds and a very short, broad, marginal cusp. Lateral teeth sickle shaped with broad base and narrow extremity.

Remarks. Orania pseudopacifica n. sp. may be compared with only three species: O. pacifica (Nakayama, 1988), with which it was confused in recent literature, O. rosea Houart, 1996 and O. pleurotomoides (Reeve, 1845). Orania pacifica (Figs 7C; 11E; 20G-H) differs in having a more strongly squamous shell with a less obvious subsutural primary cord (SP); a narrower subsutural ramp; and differently-arranged, narrower and weaker columellar folds, which are less deeply extended within the aperture (Fig. 11E). The anal notch is also narrower in O. pacifica, with a less expanded lip. Orania pacifica is widely distributed in the Indo-West Pacific, from Japan (type locality), Taiwan, the Philippines, and several other localities in the Indo-West Pacific, to Mozambique and South Africa in the Indian Ocean. Orania rosea (Fig. 20K-N), described from Reunion Island in the Indian Ocean but also found in the Philippines grows much larger, up to 23-25 mm in length, is more fusiform with a more squamous shell. The aperture is obviously larger and comparatively more strongly ovate, with a relatively smooth columellar lip except for the low abapical folds. It also has a less obvious subsutural cord. Orania pleurotomoides (Fig. 20I-J) has a more strongly shouldered shell with a less bent, narrower subsutural ramp; a narrower subsutural cord; and much narrower or absent secondary cords. In this species, the last teleoconch whorl is more sharply bent anteriorly while the siphonal canal is obviously narrower. The aperture is more triangular with a series of low abapical folds decreasing in strength abapically; a narrow, more obvious parietal tooth and a narrower anal notch with a less expanded lip. To my knowledge, none of these three species occur in French Polynesia.

Etymology. Pseudo (G) = false. Named pseudopacifica because it was misidentified as Orania pacifica in recent literature. Orania carnicolor [START_REF] Bozzetti | Muricopsis carnicolor (Gastropoda: Hypsogastropoda: Muricidae: Muricopsinae). Nuova Specie dal Madagascar Meridionale[END_REF] Type locality. South Madagascar, Lavanono, 280 kms southwest of Tolagnaro.

Other Material examined. Madagascar. ATIMO VATAE, stn TS02, 25°01.3'S, 47°00.5 'E, 18 m, 1dd;stn BB03, 25°26.4'S, 44°56.1'E, 14-18 m, 3 lv, (MNHN-IM-2009-22465, MNHN-IM-2009-22485), 1 dd; stn BB04, 25°26.9'S, 44°55.9'E, 14-18 m, 5 lv & dd; stn BS04, 25°26.9'S, 44°55.9'E, 14-18m, 1 dd; stn TS04, 25°02. 25°25.4'S, 44°54.5-7'E, 19-20 m, 6 lv (MNHN-IM-2009-22501-05, MNHN-IM-2009-22511-12); stn BP07, 25°27.2-6'S, 44°55.6-9 'E, 18-22 m, 4 dd;stn BP08, 25°27.3-6'S, 44°55.2'E, 25-26 m, 2 dd;stn BP10, 25°25.5-8'S, 44°54.4-6'E, 23-25 m, 4 lv & dd;stn TA10, 25°28.3'S, 44°55.6'E, 23 m, 1 dd;stn BS11, 25°28.6'S, 44°56.8'E, 8-11 m, 1 dd;stn BP21, 25°23.1-2'S, 44°51.4-6'E, 20-23 m (MNHN-IM-2009-22505); stn BP22, 25°23.4'S, 44°51.7 'E, 20-22 m, 5 lv & dd;stn BP31, 25°23.6-7'S, 44°53.3-5'E, 10-12 m, 1 dd;stn BP33, 25°25.80-8'S, 44°55.7-8'E, 11-13 m, 4 lv & dd;stn TA35, 24°45.6'S, 47°12.4'E, 5-6 m, 1 lv, 1 dd, (MNHN-IM-2009-22441); stn BP36, 25°21.9'S, 44°50.2 'E,[10][11][12][13][14][15][16][17]1 lv;stn BP37,19-20 m 1 lv, 1 dd; stn BP41, 25°22.9-23.2'S, 44°51.0-6 'E, 19-21 m, 3 lv;stn BP42, 25°22.8-23.7'S, 44°51.1'E, 18-21 m, 10 lv & dd;stn CP3510, 25°14.6'S, 47°09.1'E, 79-80, 1 lv (MNHN-IM-2009-22439); stn DW3519, 24°51.9'S, 47°28.0 'E,[80][81][82][83]6 lv & dd;stn CP3545,25°29'S,46°42'E,[108][109][110]1 dd;stn CP3546,25°22.7'S,46°42.5'E,[84][85]5 lv & dd;stn CP3547,25°18.0'S,46°40.3'E,[69][70] Remarks. Orania carnicolor was described as a species of Muricopsis, a genus unknown in the Indo-West Pacific, based on three dead collected specimens, but the general shape of these shells are reminiscent of several species of Orania, including O. fusulus, the type species. Its assignment to Ergalataxinae is confirmed by the morphology of the radula, here illustrated for the first time (Fig. 6E-F).

Orania castanea [START_REF] Küster | Die Gattungen Buccinum, Purpura, Concholepas und Monoceros in: Martini und Chemnitz Systematisches Conchylien-Cabinet[END_REF] Figs 6H; 21I-K Purpura castanea Küster, 1858: 170, pl. 28, figs 8, 9.

Cominella fasciata Sowerby III, 1886: 3. Cominella unifasciata var. concolor Sowerby III, 1897: 4. Cominella unifasciata nigronodulosa Turton, 1932: 53, pl. 12, fig. 302.

Thais castanea (Krauss) - [START_REF] Barnard | Contributions to the knowledge of South African marine Mollusca. Part VII. Revised fauna list[END_REF]: 691. Thais castanea -Kensley, 1973:146, fig. 507. Thais castanea (Küster, 1886) -Barnard, 1959: 224;Richards, 1981: 56, pl. 29, fig. 230;Kilburn & Rippey, 1982: 89, pl. 20, fig. 7;Steyn & Lussi, 1998: 96, Remarks. In Claremont et al (2013: 23, fig. 1;24, fig. 2), subclade Z contains "Thais" castanea, which was well supported in a cluster with Orania fischeriana (Tapparone Canefri, 1882), Orania pacifica and O. mixta Houart, 1995, and we here confirm this result (Fig 3). "Thais" castanea is clearly not a member of any rapanine or ocenebrine genus, and should be reassigned to Ergalataxinae. This assignment is further confirmed by its radula morphology (Fig. 6H), which is similar to Orania.

The date of Küster's publication of the original description of Orania castanea is wrongly indicated as 1886 by numerous authors, including one of the us (RH) and in WoRMS (MolluscaBase 2018). This is probably due to Barnard (1959: 224), who was probably the first to make this mistake. Six live specimens were collected during the recent MD 208 expedition to the Walters Shoal Mountain (Fig. 21K). This range extension, even if not unexpected in this area, remains very interesting.

Orania pachyrhaphe [START_REF] Smith | On a collection of Mollusca from Japan[END_REF] Fig. 22A-E Fusus pachyrhaphe E.A. [START_REF] Smith | On a collection of Mollusca from Japan[END_REF]: 205, pl. 20, figs 37, 37a. Bedeva bireleffi -Tsuchiya, 2000: 381, pl. 189, fig. 90 [not Bedevina birileffi (Lischke, 1871)]. 1871), the intricate history of which was analysed by Houart et al. (2013). The smallest of the four paralectotypes is a young specimen of O. pachyrhaphe (Fig. 22C). A specimen of B. birileffi is here illustrated (Fig. 22H-I) together with the paralectotype figured by E.A. Smith (1879: fig. 37a) (Fig. 22F-G).

Bedevina was analysed in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] and its validity in the Ergalataxinae was confirmed. However, the boundaries of this genus remain unclear because the same clade contained some species included in Spinidrupa (but not the type species). The definition of Bedevina and Spinidrupa thus remain to be delimited exactly, but, as noted in Claremont et al. (2013: 27), the conservative retention of both genera is recommended until further analyses are undertaken. One species, P. palmeri [START_REF] Powell | New Zealand Molluscan systematics with descriptions of new species, part 6[END_REF]) (Fig. 22O-P) from New Zealand, is here added as a new combination. In the original description by Powell (1967: 193) Radula of the type species (Fig. 8C) with three dimensional rachidian tooth bearing a long, moderately broad, projecting central cusp, a very small, narrow lateral denticle, a moderately long, broad, lateral cusp and strong marginal folds giving rise to small, short marginal denticles, and a bifid marginal cusp. Lateral teeth sickle shaped, with broad base.

Figure 22

A-E. Orania pachyrhaphe (E.A. Smith, 1879) A-B. Japan, Ukushima, Goto Islands, 33°15.5' N, 129°5 ' E, 11 (Dall, 1908), Easter Id, Hanga Piko, RH, 13.8 mm. M-N. Pascula darrosensis (E.A. Smith, 1884), Philippines, Sulu Sea, RH, 10.3 mm. O-P. Pascula palmeri [START_REF] Powell | New Zealand Molluscan systematics with descriptions of new species, part 6[END_REF], New Zealand, Poor Knights Islands, on rock wall, 20 m, RH, 20.6 mm.

Remarks.

The genus Tenguella currently includes five Recent species: Tengualla ceylonica (Dall, 1923), T. granulata (Duclos, 1832), T. hoffmani Houart, 2017, T. marginalba [START_REF] Blainville | Disposition méthodique des espèces récentes et fossiles des genres Pourpre, Ricinule, Licorne et Concholepas de M. de Lamarck, et description des espèces nouvelles ou peu connues, faisant partie de la collection du Muséum d'Histoire Naturelle de Paris[END_REF] and T. musiva [START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF]. Two new species are described below giving a new total of 7 species in this genus (Table 2). Type localities. Purpura granulata: Australia ("Nouvelle-Hollande"); Purpura tuberculata: Red Sea and Madagascar, here restricted to the Red Sea; Purpura tuberculata var. cingulifera: unknown.

Distribution. Tenguella granulata is a very common species living throughout the Indo-West Pacific.

Remarks. Tenguella granulata is here compared with two species, T. chinoi n. sp. and T. ericius n. sp., collected during the MNHN/IRD expeditions in The Marquesas, Vanuatu and Papua New Guinea and known from a few other localities (see below). Other material examined. Papua New Guinea: PAPUA NIUGINI, stn PFM03, Bilbil Island, exposed steep, fringing reef, under rocks, exposed, 05°15'S, 145°47'E, 2 m, 2 lv (syntopic with Tenguella granulata); stn PM25, Barag Island, 05°01'S, 145°48'E, fringing reef, on narrow barrier island, 3 lv; stn PM33, Rempi area, bay on south side, coral reef, 05°02'S, 145°48'E, coral reef, 1 lv; stn PM43, Wonad Island, 05°08'S, 145°49'E, night tide, sandy beach and intertidal rocks, 0-1 m, 1 lv (syntopic with Tenguella granulata); stn PR76, Rempi area, SW Hargun Island, 05°02'S, 145°48'E, 2-15 m, 1 lv (syntopic with Tenguella granulata); stn PR192, Kranket Island, Cape Jantzen, 05°12'S, 145°49'E, 3-60 m, 2 lv; stn PR204, BilBil I., southern tip, no coordinates, 3-15 m, 2 lv (syntopic with Tenguella granulata); stn PR214, Tab Island, no coordinates, 1-8 m, 4 lv & 1 dd (syntopic with Tenguella granulata and T. ericius n. sp.); Hansa Bay, Madang Province, Laing Island, 2 lv, juv, (RH).

China Sea: (no other data), 11 lv (ex Staadt coll., 1969) (MNHN). Western Australia: Exmouth, 1 lv (RH).

Distribution. Western Australia, Exmouth (1 RH), East Java [START_REF] Dharma | Recent and fossil Indonesian shells[END_REF], Japan, Izu Islands and southwards [START_REF] Tsuchiya | Muricidae[END_REF], Papua New Guinea and Vanuatu; perhaps also the China Sea (MNHN). Intertidal to 3 m depth, sandy beach and intertidal rocks (Fig. 24).

Description. Shell small for the genus, up to 13.6 mm in length at maturity (paratype MNHN-IM-2000-34216) (Fig. 25B-C). Length/width ratio 1.3-1.4. Broad, heavy, nodose. Subsutural ramp broad, weakly sloping, concave. Blackish-brown or black. Aperture white within with narrow black coloured narrow band at edge and dark bluish broad blotch at adapical part of strong curve of columellar lip. Spire high, low. Teleoconch of up to 5 or 6 strongly convex, shouldered, nodose whorls. Suture adpressed. Protoconch and early teleoconch whorls strongly eroded in type material. Axial sculpture of teleoconch whorls consisting of low, broad, nodose ribs with strong, low, broad nodes when crossing primary spiral cords. Last teleoconch whorl with 8 or 9 low, occasionally indistinct ribs, eroded on earlier whorls. Spiral sculpture of low, strong, rounded and flat, broad, nodose, primary cords and low, narrow threads. Last teleoconch whorl with SP, P1-P5 with 2-4 flat, narrow, smooth threads between each pair of cords. SP very broad, obvious, usually broader than other spiral cords. P1-P2 similar in size, P3-P5 decreasing in strength abapically. P5 very narrow. Aperture large, very narrow, strongly ovate, sinuous. Columellar lip broad, strongly curved, with one or two very narrow knobs abapically and low parietal tooth at adapical extremity. Rim weakly partially erect abapically, adherent adapically. Anal notch narrow, moderately deep. Outer lip weakly erect, smooth, edge broad with 4 strong, high denticles within, consisting of D1-D4. D1 very broad and high, D2 more than half the size of D1, D3 and D4 lower and narrower, more elongate within aperture. Siphonal canal very short, 9-10% of total shell length, narrow, straight, broadly open. Radula consisting of a rachidian with a moderately large, long, central cusp, a short, narrow, lateral denticle and a moderately broad, long, lateral cusp on each side, 2 or 3 obvious, short, marginal denticles and a short marginal cusp. Lateral teeth sickle shaped with broad base and narrow extremity.

Remarks. Based on molecular data, Tenguella chinoi n. sp. was separated from Tenguella granulata, T. ceylonica, T. musiva and T. marginalba by Claremont et al (2013, as "Tenguella n. sp."). These five species formed a well-supported subclade. We here confirm this result (Fig 3). The material studied in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] (as Tenguella n. sp.) consists of six specimens, four of them were crashed to obtain the soft parts, two remained intact, one of those is here illustrated (Fig. 25P). These two intact specimens are obviously what is here described as T. ericius n. sp. and so, the NHMUK material was first identified as this species. However, in the meantime, two specimens of T. ericius n. sp. from the PAKAIHI I TE MOANA expedition in the Marquesas Archipelago were also analyzed in MNHN and proved to be different with around 10% of genetic distance (COI gene) from the specimen analyzed in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF]. This unexpected result could mean two things: either the Marquesas species is a sibling species but is genetically different, or the material in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] was composed of different species and the specimen analyzed by these authors was not T. ericius n. sp. After a very careful examination of the crushed shells, the reconstruction of the aperture of one studied specimen was possible (Fig. 25A) and it is clearly this second hypothesis which is the correct one. The species studied in Claremont et al. ( 2013) is clearly the species described here as T. chinoi n. sp. while the two intact, not studied specimens are undoubtedly T. ericius n. sp. These two species, T. ericius n. sp. and T. chinoi n. sp. are syntopic in some localities and thus also in Guam, where the specimens analyzed in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] were collected. Thanks now to the results obtained by the analysis of T. ericius n. sp. from the Marquesas it is also proved that the two species are genetically different. For other differences between these two species see under T. ericius n. sp. The protoconch and the first teleoconch whorls were preserved in two juvenile specimens from Papua New Guinea (RH). The protoconch is conical and glossy brown, consisting of 3.5 whorls and a terminal lip of the sinusigera type (Fig. 25G). Some specimens collected in the China Sea (no other locality data) before 1960 and deposited in MNHN are bigger and reach a length of 16 mm (Fig. 25H). The shell morphology of this little species confirms its separation from the other Tenguella species and, more particularly, from the similar looking T. granulata. Tenguella granulata is a species with a quite variable shell morphology (Fig. 25Q-S), but it differs consistently from T. chinoi n. sp. in being less stocky, with more acute nodes, in having a comparatively larger shell, often reaching a length of more than 25 mm, with an average size of 15-20 mm, compared to the 12-16 mm of T. chinoi. The spire in T. granulata is higher and more acute; the aperture is larger, broader, and less sinuous, with relatively smaller, less obvious denticles and a narrower outer apertural lip. Tenguella chinoi n. sp. was illustrated by [START_REF] Dharma | Recent and fossil Indonesian shells[END_REF] as Azumamorula sp. This monotypic genus still needs to be analysed carefully, but A. mutica (Lamarck, 1816), the type species of Azumamorula (Fig. 27A-B), differs consistently from Tenguella in having a broadly convex, smoother shell with obsolete axial ribs and very low spiral sculpture. It also has a broader aperture with comparatively weaker denticles.

Etymology. Named after Mitsuo Chino (Japan), known for his numerous contributions to malacology and author of several new species, who sent me some of the shells included in the studies of Ergalataxinae. Description. Shell small for the genus, up to 15.3 mm in length at maturity. Length/width ratio 1.4-1.5. Biconical, broad, heavy, weakly spinose and nodose, tuberculate. Subsutural ramp strongly sloping, concave. Blackish-brown or black. Aperture white within with narrow black coloured narrow band at edge and dark bluish broad blotch at adapical weak curve of columellar lip. Spire moderately high. Teleoconch of up to 5 broad, shouldered, nodose whorls. Suture adpressed. Protoconch eroded in all specimens. Axial sculpture of teleoconch whorls consisting of narrow, moderately high, nodose ribs. Early spire whorls with 12 or 13 ribs; last whorl with 9 ribs with strong, relatively high, broad nodes at intersection with primary spiral cords. Nodes higher and more apparent on apertural rib. Spiral sculpture of low, strong, rounded, broad, nodose, primary cords with 2 or 3 very narrow, rounded threads between each pair of cords. Last teleoconch whorl with broad SP, similar to P1-P3, P4 narrower, P5 narrowest primary cord. Aperture large, ovate. Columellar lip broad, smooth or with one or two very small denticles abapically and low parietal tooth at adapical extremity. Anal notch moderately deep, broad, with an open channel shortly extending in SP spine. Outer lip weakly erect, smooth, with 4 strong, broad denticles within consisting of D1-D4; D1 broadest and strongest, D2-D4 decreasing in strength abapically. Siphonal canal very short, 3.5-5.5% of total shell length, narrow, straight, broadly open. Operculum and radula unknown.

Remarks. Tenguella chinoi n. sp. differs consistently from T. ericius in having a slightly broader, more globose, stocky and solid shell, with broader and lower axial ribs; and in having blunt, broad nodes compared to the more spiny shell of T. ericius n. sp. T. chinoi also has a more sinuous columellar lip and broader, more apparent apertural denticles within the outer lip, resulting in a much more narrower aperture as opposed to a relatively broad aperture in T. ericius n. sp. In T. chinoi the spire is also lower and less acute, the outer apertural lip is relatively broader, and the siphonal canal, although short in T. chinoi n. sp., is comparatively longer (9-10% of the total shell length) than in T. ericius (3.5-5.5 % of the shell length). Tenguella granulata differs from T. ericius n. sp. in having an obviously larger shell and a comparatively narrower aperture due to a strongly folded columellar lip. The outer apertural lip of T. granulata is almost similar than that of T. ericius n. sp., but it is obviously less expanded abaperturally at the level of P4, between D3 and D4 (Fig. 12D).

Etymology. Ericius: hedgehog, named after its spinose shell. Since then, we have had the opportunity to examine more specimens of Claremontiella species from off West Africa, especially a population living in São Tomé (Fig. 27O-P). In these specimens, the shell looks strongly similar to the original illustration of Cantharus consanguineus. Having sent typical specimens of C. nodulosa from Virgin Islands, close to Jamaica, the type locality of C. nodulosa and a species identified as C. consanguinea from São Tomé for genetic analyses, we received an opinion from Andrea Barco (in litt, 17 June 2013) who said, "There is little genetic distance between the two groups, but it is really low. I would expect higher values between different species." There is little doubt that some genetic differences could exist between populations from São Tomé and Saint Helena, so the names Claremontiella consanguinea and C. nodulosa could be subjective synonyms. Further analyses are needed before making a final decision. In the meantime C. consanguinea is here provisionally retained as a valid species. Claremontiella adiakritos n. sp. is assigned to Claremontiella because of its similar shell and radula morphology. The shell of C. adiakritos is close to the type species (Figs 27H and28E) with which it has often been confused, as Morula, Evokesia, or Pascula ferruginosa (see under C. adiakritos n. sp.).

Discussion

Etymology. The new genus is named for Martine Claremont whose contribution to the molecular phylogeny of Muricidae, in particular here for the Ergalataxinae, while working at the Natural History Museum, London, was extremely useful and appreciated. Distribution. Magdalena Bay, Baja California, through the Gulf of California and south along the Sonoran coast of Mexico at least as far as Guaymas, intertidally, under rocks [START_REF] Keen | Sea shells of Tropical West America, Marine Mollusks from Baja California to Peru. 2d edit[END_REF]. Its distribution south to Bocapán, El Rubio and Tumbes, Peru [START_REF] Alamo | Lista sistemática de moluscos marinos del Perú[END_REF][START_REF] Houart | Les Muricidae d'Afrique Occidentale -II. Ocenebrinae, Ergalataxinae, Tripterotyphinae, Typhinae, Trophoninae & Rapaninae[END_REF][START_REF] Skoglund | Panamic Province Molluscan Literature -Additions and Changes From 1971 through 2001 -III. Gastropoda[END_REF] is doubtful (see remarks). Finet (1994: 50) mentions records from the Galapagos Islands but without any illustration, which is also here considered doubtful (Fig. 30).

Description. Shell large for the genus, up to 24.7 mm in length at maturity (paratype RH). Length/width ratio 2.1-2. The ontogeny of the spiral cords is unknown since all examined specimens were badly corroded from the first to the second or third teleoconch whorls. For this reason, the protoconch could also not be examined. However, Radwin & D'Attilio (1972: 338) described and illustrated the protoconch of C. adiakritos n. sp.

(as Evokesia ferruginosa) with 3.5 papillose whorls, similar to the protoconch morphology of the type species of Claremontiella (Fig. 27J).

The radula of this species, illustrated in Radwin & D'Attilio (1972: 340, fig. 16), is strangely different from that illustrated here (Fig. 8D), extracted from a specimen collected near San Felipe. The drawing shows a broader and shorter central cusp, and broader lateral cusps as well as broad lateral teeth, while the SEM here illustrated shows a typical ergalataxine radula, very close to that of C. consanguinea, with a narrow, long central cusp, a very short lateral denticle on each side, short, moderately broad lateral cusps and narrow, slender, lateral teeth. The assignation of this species to Claremontiella n. gen. is tentative and requires genetic analyses for confirmation. [START_REF] Alamo | Lista sistemática de moluscos marinos del Perú[END_REF][START_REF] Houart | Les Muricidae d'Afrique Occidentale -II. Ocenebrinae, Ergalataxinae, Tripterotyphinae, Typhinae, Trophoninae & Rapaninae[END_REF] Other included species. Murichorda rumphiusi (Houart, 1996) Distribution. Throughout the Indo-Pacific.

Remarks. Murichorda fiscellum was previously assigned to Cronia H. & A. Adams, 1853 by Cernohorsky (1969: 311;1982: 113), but Cronia, type species C. amygdala [START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF], was included in a different clade by [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF]), together with Ergalatax Iredale, 1931and Maculotriton Dall, 1904, which could be potential synonyms (see Table 2).

Because its shell morphology is close to M. fenestrata [START_REF] Blainville | Disposition méthodique des espèces récentes et fossiles des genres Pourpre, Ricinule, Licorne et Concholepas de M. de Lamarck, et description des espèces nouvelles ou peu connues, faisant partie de la collection du Muséum d'Histoire Naturelle de Paris[END_REF], the type species of Muricodrupa, Murichorda fiscellum has been assigned to Muricodrupa by numerous authors, starting with [START_REF] Kay | Hawaiian Marine Shells. Reef and shore fauna of Hawaii[END_REF]. However, in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] "Morula" fiscella and "Morula" rumphiusi formed a wellsupported clade in all analyses but were excluded from Muricodrupa. Both are therefore here assigned to Murichorda new genus, along with Murichorda jacobsini. (Reeve, 1845)

Etymology. Prefix

Figure 1 .

 1 Figure 1. Method for determining diameter, height and counting the number of protoconch whorls. Terminology used to describe the spiral cords and the apertural denticles (after Merle 2001, 2005) (Figs 10-12). Variable features are given in parentheses.

  t: tertiary cord Aperture D1 to D5: abapical denticles; ID: infrasutural denticle.

Figure 2 .

 2 Figure 2. Terminology used to describe the radula, here Pascula darrosensis (E.A. Smith, 1884) cc: central cusp; ld: lateral denticle; lc: lateral cusp; ma: marginal area; LT: lateral teeth.

Figure 3 .

 3 Figure 3. Bayesian phylogeny of Ergalataxinae included in this paper and rapanine outgroups

  Fig. 6A) with three dimensional rachidian tooth bearing a long, narrow, projecting central cusp, a small, very narrow lateral denticle, a moderately long, narrow, lateral cusp and weak marginal folds. No marginal cusp. Lateral teeth sickle shaped, with broad base. Remarks. The genus Cytharomorula includes 10 Recent species in WoRMS (MolluscaBase 2018): Cytharomorula ambonensis (Houart, 1996) (Indo-West Pacific); C. benedicta (Melvill & Standen, 1895) (Pacific Ocean); C. danigoi Houart, 1995 (Pacific Ocean); C. dollfusi (Lamy, 1938) (Indo-West Pacific); C. grayi (Dall, 1889) (western and eastern Atlantic, southwestern Indian Ocean); C. lefevreiana (Tapparone Canefri, 1880) (western Indian Ocean); C. paucimaculata (Sowerby, 1903) (Japan); C. pinguis Houart, 1995 (New Caledonia, New Hebrides Arc and Tonga); C. springsteeni Houart, 1995 (Indo-West Pacific); C. vexillum

Figure 5 .

 5 Figure 5. Distribution of Cytharomorula arta n. sp.

Figure 6 Figure 7 Figure 8 Figure 9

 6789 Figure 6 -Radulae A. Cytharomorula vexillum Kuroda, 1953, New Caledonia. Scale bar: 50 µm (SEM A. Warén) B-C. Cytharomorula elegantula n. sp., New Caledonia. Scale bars: B. 100 µm; C. 20 µm (SEM P. Bouchet) D. Cytharomorula ornamentata (Houart, 1995), Transkei, Sourth Africa. Scale bar: 50 µm (SEM A. Warén) E-F. Orania carnicolor Bozzetti, 2009, Madagascar. Scale bars: E: 50 µm; F. 20 µm (SEM A. Warén) G. Orania fusulus (Brocchi, 1814), Angola. Scale bar: 50 µm (SEM A. Warén) H. Orania castanea (Küster, 1858), East London, South Africa. Scale bar: 50 µm (SEM A. Warén)

Figure 10 -

 10 Figure 10 -Spiral sculpture and apertural morphology A-B. Cytharomorula arta n. sp., Reunion, MD32, stn DC56, 21°05' S, 55°12' E, 1982, 170-225 m. A. Paratype MNHN-IM-2000-34205, 12.3 mm; B. Holotype MNHN-IM-2000-34203, 14.3 mm. C. Cytharomorula elegantula n. sp., New Caledonia, Kaimon Maru Bank, SMIB 8, stn DW159, 28/01/1993, 24°46' S, 168°08' E, 241-245 m, paratype MNHN-IM-2000-34209, 15.7 mm. D. Cytharomorula grayi (Dall, 1889), Canary Islands, Tenerife, Palma, in shrimp nets, 150-200 m, 1985, RH, 19.6 mm. E. Cytharomorula fatuhivaensis n. sp., French Polynesia, Marquesas Islands, Fatu Hiva, 03/09/1990, 10°31' S, 138°39' W, 210 m, holotype MNHN-IM-2012-18829, 14.7 mm. F. Cytharomorula springsteeni Houart, 1995, Philippines, tangle nets, 146 m, paratype RH, 12.7 mm.

Figure 11 -

 11 Figure 11 -Spiral sculpture and apertural morphology A-B. Cytharomorula grayi (Dall, 1889), MD208, stn DW4877, Walters Shoal, 33°10'S, 43°49'E, 217-256 m, MNHN-IM-2013-66419, 23.2 mm. C-D. Orania pseudopacifica n. sp., French Polynesia, Marquesas Islands, Nuku Hiva, 25/08/1997, 8°45' S, 140°13' W, 104-109 m, holotype MNHN-IM-2000-34211, 14.5 mm; D. Detail of the columella. Scale bar: 1 mm. E. Orania pacifica (Nakayama, 1988), Philippines, Panglao, 55-81 m, detail of the columella, RH. Scale bar: 1 mm. F. Cytharomorula absidata n. sp., New Caledonia, NORFOLK 2, stn DW2123, 23°18'S, 168°15'E, 187-197 m, Holotype MNHN-IM-2000-34201, 9.6 mm. G. Claremontiella adiakritos n. sp., West Mexico, Mazatlan, holotype NHMUK 20180545, 23.3 mm.

Figure 12 -

 12 Figure 12 -Spiral sculpture and apertural morphology A-B. Tenguella chinoi n. sp., Papua New Guinea, Papua Niugini, stn PR214, 08/12/2012, 1-8 m (no other information), MNHN, 12.5 mm. C-D. Tenguella ericius n. sp., Vanuatu, SANTO, stn FM36, 15°22,4'S 167°13'E 27/09/2006 0-1 m, paratype MNHN-IM-2000-34218, 13.9 mm.

Figure 17 .

 17 Figure 17. Distribution of Cytharomorula manusuduirauti n. sp. Remarks. Cytharomorula manusuduirauti n. sp. has been confused with C. pinguis Houart, 1995 and C. springsteeni Houart, 1995. However both species differ from C. manusuduirauti. Cytharomorula pinguis (Fig. 18M-N) differs in having a comparatively narrower shell with broader, flatter and less numerous spiral cords. C. pinguis also has a comparatively narrower aperture with strong D1 denticle and strong, heavy knobs on the columellar lip, as opposed to the smooth columellar lip with very weak adapical knobs in C. manusuduirati n. sp. Both species have primary, secondary and tertiary cords of approximately similar strength, which are difficult to separate. However, C. pinguis has 20-22 broad, flat cords on the last teleoconch whorl as opposed to 37-40 narrow, rounded cords in C. manusuduirauti n. sp.

Figure 19 .

 19 Figure 19. Distribution of Orania pseudopacifica n. sp.

  comb. nov. Figs 6E-F; 21D-H Muricopsis carnicolor Bozzetti, 2009: 19, text figs. Type material. Holotype MNHN-IM-2000-22900.

Figure 20 (

 20 Figure 20 (scale bar: 500 µm) A-F. Orania pseudopacifica n. sp. French Polynesia, Marquesas Islands, Nuku Hiva, 8°45' S, 140°13' W, 104-109 m. A-D. Holotype MNHN-IM-2000-34211, 14.5 mm; E-F. Paratype MNHN-IM-2000-34212, 14.0 mm.G-H. Orania pacifica(Nakayama, 1988), Philippines, Balut Island, RH, 17.5 mm. I-J. Orania pleurotomoides(Reeve, 1845), Philippines, Mactan Island, 60 m, RH, 16.1 mm. K-N. Orania rosea Houart, 1996 K-L. Reunion Island, off Saint Pierre, 21°21'S 55°27'E, 73-77 m, holotype MNHN-IM-2000-910, 18.6 mm, photo Manuel Caballer (MNHN) E-Recolnat Project: ANR-11-INBS-0004; M-N. Philippines Mactan Id, Cebu, 120 m, sand and broken coral, RH, 20.9 mm.

Fig. 23 .

 23 Fig. 23. Purpura granulata. Original illustration from Duclos, 1832.

Figure 24 .

 24 Figure 24. Distribution of Tenguella chinoi n. sp. Tenguella ericius n. sp. Figs 12C-D; 25I-P; 26 Morula granulata -Wells et al., 1990: 44, pl. 21, fig. 141 (not Tenguella granulata).

Figure 26 .

 26 Figure 26. Distribution of Tenguella ericius n. sp.

Figure 30 .

 30 Figure 30. Distribution of Claremontiella adiakritos n. sp. Murichorda new genus Figs 8G-H; 9A; 28F-L Type species: Purpura fiscellum Gmelin, 1791: 3552, Indo-West Pacific.

  Trophon lowei[START_REF] Watson | On the marine Mollusca of Madeira; with descriptions of thirty-five new species, and an index-list of all the known seadwelling species of that island[END_REF], Madeira, lectotype NHMUK 1911.7.17.2, 19.3 mm, photo Harry Taylor, NHMUK Photographic Unit, © Natural History Museum of London; O-P. Cantharus laevis E.A.[START_REF] Smith | Report on the marine fauna of the Island of St. Helena[END_REF], St Helena, lectotype NHMUK 1889.10.1.2362, 22.2 mm, photo Harry Taylor, NHMUK Photographic Unit, © Natural History Museum of London.Remarks. Cytharomorula elegantula n. sp. from New Caledonia (Fig.16H-N), also described herein, differs from C. fatuhivaensis n. sp. in having a usually lighter, narrower shell, with a length/width ratio of 2.0-2.1 as opposed to the 1.8-2.0 of C. fatuhivaensis (C. fatuhivaensis is stockier and broader); in having a narrower and longer siphonal canal, 16-18% of total shell length, vs 12-14 % in C. fatuhivaensis; a higher, more elongate spire; narrower, lower, usually more numerous axial ribs; a comparatively more elongate aperture; less apparent, usually narrower spiral cords, except for P4 which is very often higher and more apparent in C. elegantula than has been observed in other Cytharomorula species, but which is of equal height to other cords in C. fatuhivaensis.

	18% of total shell length, narrow, dorsally recurved,	Cytharomorula grayi -Houart & Tröndlé, 2008: 70, Type locality. Philippines, Aliguay Island, near 8°44'
	broadly open.	92 [(not C. grayi (Dall, 1889)]. N, 123°12'40'E, sand, sandy gravel & sandy mud
	Operculum unknown. Radula with three dimensional	bottoms, 50-150 m.
	rachidian tooth bearing a long, narrow, projecting	Type material. Holotype MNHN-IM-2012-18829,
	central cusp, a small, narrow lateral denticle on each	French Polynesia, Marquesas, Fatu Hiva, SMCB: stn Distribution. Philippines Islands, living at 150-160 m
	side, a medium sized lateral cusp, as broad as central	CAS303, 10°31'S, 138°39'W, 210 m, lv. and Japan, Ogasawara Islands (Tsuchiya, 2017) (Fig.
	cusp but shorter, and a few marginal folds. No	Paratypes: French Polynesia, Marquesas, Fatu Hiva, 17).
	marginal cusps. Lateral teeth sickle shaped, relatively	10°31'S, 138°39'W, 210 m, 16 lv, MNHN-IM-2000-
	broad.	34210, 1 lv, RH. Description. Shell small, up to 11.6 mm in length
		(holotype). Length/width ratio 1.9-2.0. Biconical,
	Remarks. Cytharomorula grayi, with which C. elegantula n. sp. has often been confused, differs in having a more strongly shouldered, less fusiform shell	broadly ovate, heavy, lightly nodose. Type locality. French Polynesia, Marquesas, Fatu Light tan, extreme abapical part of last teleoconch Hiva, 10°31'S, 138°39'W, living at 210 m. whorl and siphonal canal dark brown or blackish
	and tertiary cords. First teleoconch whorl with visible SP, IP, P1 and P2; second whorl with SP, adis, IP, P1, with a narrower, more concave and less strongly sloped subsutural ramp, and broader, stronger, higher axial ribs, broader and more obvious on shoulder. These axial ribs are also less numerous, 6 or 7 on the last teleoconch whorl in C. grayi as opposed to 8 or 9 in C. elegantula n. sp. C. grayi also has a broader, less fusiform aperture with more obvious denticles. F-L. Cytharomorula absidata n. sp. F-H. New Caledonia, NORFOLK 2, stn DW2123, 23°18'S, 168°15'E, 187-197 m, holotype MNHN-IM-2000-Aperture narrow, ovate. Columellar lip narrow with weak, low parietal tooth at adapical extremity and 2 or brown. Occasionally with weak brown blotches Other material examined. MUSORSTOM 9, stn between axial ribs of last teleoconch whorl. Aperture DW1145, Marquesas, Ua Pou, 9°19'S, 140°06'W, 150-white, columellar lip occasionally with brown blotch 180 m, MNHN-IM-2008-2929, 7 lv & dd; stn DW1146, 9°19'S, 140°06'W, 200 m, MNHN-IM-2008-2928, 3 dd; stn DW1148, 9°19'S, 140°06'W, 300 m, MNHN-IM-2008-2927, 2 dd; stn DW1197, Hiva Oa, 9°57'S, 140°02'W, 277-372 m, MNHN-IM2008-2926, 1 dd; stn DR1247, Fatu Hiva, 10°34'S, 138°42'W, 1150-1250 m, MNHN-IM-2008-2925, 1 dd. 34201, 9.6 mm; I. New Caledonia, NORFOLK 2, stn DW2123, 23°18'S, 168°15'E, 187-197 m, paratype MNHN-IM-2007-18225 (BOLD MUBA765-18; GenBank HE584051.1) (fragment of analysed specimen, 8 mm); J-L.New Caledonia, SMIB 5, stn DW93, 22°20'S, 168°43'E, 240-255 m, paratype MNHN-IM-2000-34202, 13.7 mm. A-B. Reunion, MD32, stn CP57, 21°05' S, 55°11' E, 1982, 210-227 m, holotype MNHN-IM-2000-34203, 14.3 mm; C-D. Reunion, MD32, stn CP57, 21°05' S, 55°11' E, 1982, 210-227 m, paratype MNHN-IM-2000-34206, 11.9 mm; E. Protoconch, Reunion, MD32, stn DC2, 1982, 21°12' S, 55°49' E, 160-190 m, paratype MNHN-IM-2000-34204. F-G. Cytharomorula lefevreiana (Tapparone Canefri, 1880), MD32, Stn DC85, Reunion, 20°59' S, 55°15' E, 58-70 m, MNHN, 6.6 mm. H-P. Cytharomorula grayi (Dall, 1889) H-I. Canary Islands, Tenerife, Palma, in shrimp nets, 150-200 m, RH, 19.6 mm; J. East of Guadeloupe, south of Desirade, 250 m, RH, 13.8 mm; K. West coast of Barbados, dredged 165 m, RH, 13.2 mm; L South-Eastern Brazil, Martin Vaz Id, 20°32' S, 28°52' W, 780-795 m (dd), MNHN-IM-2012-33133, 14.1 mm; M. Madeira, RH, locality. Cytharomorula manusuduirauti n. sp. Figs 17; 18G-L Cytharomorula pinguis -Tsuchiya, 2000: 381, pl. 189, fig. 88; Houart, 2008: 23, 198, pl. 394, fig. 6 [not Type material. Holotype HMNS 2017.2223 Po.406866, Philippines, Aliguay Island, near 8°44' N, 123°12'E, sand, sandy gravel & sandy mud bottoms, 50-150 m, lv. Paratypes: Philippines, Balicasag Island, near 9°31' N, 120°41'E, tangle or lumun-lumun net, 100-150 m, lv, 1 HMNS 2017.2223 Po.524523; Philippines, Balicasag Island, near 9°31' N, 120°41'E, tangle or lumun-lumun net, 50-80 m, dd, 1 HMNS 2017.2223 Po.571785; Philippines, Aliguay Island, rocky mud, 160 m, lv, 1 RH. abapically. Spire high with 3+ protoconch whorls (first whorl broken) and teleoconch of 4 or 4.5 broad, weakly shouldered, nodose whorls, spire whorls more strongly shouldered. Suture adpressed. Protoconch large, high, conical, glossy. Maximum width 800 µm, height 1000 µm. Terminal lip of sinusigera type. Axial sculpture of teleoconch whorls consisting of moderately high, broad, rounded ribs, with 8 or 9 ribs on all whorls, ribs obviously broader on last whorl. Spiral sculpture of numerous, narrow, rounded cords, more than 35 on last whorl, equally distant and of same strength on whole length of shell. Aperture small, ovate. Columellar lip narrow, smooth or with two weak, narrow knobs abapically. Rim weakly erect abapically, adherent adapically. Anal notch deep, broad. Outer lip smooth, with 6 strong, elongate denticles within, decreasing in strength abapically, consisting of ID, D1-D5. Siphonal canal very short, 8-10% of total shell length, narrow, weakly dorsally recurved, broadly open. Operculum and radula unknown. Figure 16 (scale bars: 500 µm) A-G. Cytharomorula grayi (Dall, 1889) A-D. MD208, stn DW4877, Walters Shoal, 33°10'S, 43°49'E, 217-256 m; A-C. MNHN-IM-2013-66419 (BOLD MUBA781-18; GenBank MK216540), 23.2 mm; D. MNHN-IM-2013-66418, 22.0 mm; E. Protoconch, Walters Shoal, MD208, stn DW4894, 33°09'S, 43°50'E, 199-261 m, MNHN; F. South Africa, Durban, off Umlaas Canal, coarse rubble, 14/12/1983, 128 m, NM B6282, 19.2 mm; G. SW Grande Glorieuse Id, BENTHEDI, stn DS93, 07/04/1977, 11°33' S, 47°16' E, 480-550 m, MNHN, 17.2 mm. H-N. Cytharomorula elegantula n. sp. 17.8 mm; N. Etymology. Named after Fatu Hiva Island, the type H-J. New Caledonia, KANACONO, stn DW4778, 23°03'S, 168°18'E, 170-248 m, holotype MNHN-IM-2013-
	s1, P2, s2; third and fourth with additional abis and 63326, (BOLD MUBA780-18; GenBank MK216542), 10.2 mm; K-L. New Caledonia, N.O. "Jean Charcot" 3 elongate, low, narrow folds abapically. Rim weakly
	tertiary cords. Last whorl with SP, adis, IP, abis, P1,	partially erect, adherent at small portion at adapical
	s1, P2, s2, P3, s3, P4, s4, P5, s5, P6, s6, ADP, ads, MP	extremity. Anal notch deep, broad. Outer lip erect,
	and ABP; P4 occasionally broader and higher with	smooth with 6 weak, low, elongate denticles within,
	high knobs at intersection with axial ribs, then usually	consisting of ID, D1-D5. Siphonal canal short, 16-
	lighter or white coloured.

Tenguella ericius n. sp., Vanuatu, SANTO, stn FM36, 15°22,4'S 167°13'E 27/09/2006 0-1 m, paratype

MNHN-IM-2000-34218, 13

.9 mm. For differences with Cytharomorula absidata n. sp., C. arta n. sp. and C. fatuhivaensis n. sp. see the descriptions of these species herein. The Pleistocene specimens from Vanuatu were collected during an exploration in February 2006 of fossil sites as part of the SANTO 2006 Expedition

[START_REF] Lozouet | The Holocene and Pleistocene marine faunas reconsidered[END_REF]

. The fossil specimens come from the Kere 1 deposit (= Kere Shellbed of Mallik and Ladd). A well-preserved coral (Flabellum sp.) collected in this outcrop gave an age of 133 000 yr.

[START_REF] Lozouet | The Holocene and Pleistocene marine faunas reconsidered[END_REF]

(see also

Houart, 2013b)

.

Etymology. Elegantula (L): diminutive of elegans, meaning small and elegant.

Cytharomorula fatuhivaensis n. sp.

Figs 10E; 14; 18A-F Cytharomorula sp. cf. C. grayi

[START_REF] Tröndlé | Les Muricidae de Polynésie Française[END_REF]: 88, fig. 48 [(not C. grayi (Dall, 1889)

].

Distribution. French Polynesia, Fatu Hiva, Ua Pou and Hiva Oa, living at 180-210 m (Fig.

14

).

Description. Shell medium sized for the genus, up to 16.5 mm in length at maturity (paratype MNHN).

Length width ratio 1.8-2.0. Biconical, broadly ovate, heavy, nodose. Subsutural ramp broad, strongly sloping, weakly concave. Light tan or whitish with dark brown spiral cords. Aperture white. Spire high with 3.5 protoconch whorls and teleoconch up to 5 broad, strongly convex, weakly shouldered, nodose whorls. Suture strongly adpressed. Protoconch small, conical. Whorls smooth, glossy, with a very narrow keel abapically. Height and width 1000 µm. Terminal lip erect, of sinusigera type. Figure 13 (scale bar: 500 µm) A-C. Cytharomorula vexillum Kuroda, 1953. A-B. Japan, Tosa, 128 m, MNHN, 21.1 mm; C. Protoconch, New Caledonia. D-E. Cytharomorula ornamentata (Houart, 1995), South Africa, Durban, RH, 16.8 mm. M-O. Cytharomorula springsteeni Houart, 1995 M. Philippines, tangle nets, 146 m, paratype RH, 12.7 mm; N-O. New Caledonia, Kaimon Maru Bank, SMIB 8, stn DW159, 28/01/1993, 24°46' S, 168°08' E, 241-245 m, 13.2 mm, MNHN. Axial sculpture of teleoconch whorls consisting of high, strong, broad, rounded, nodose ribs. First and second teleoconch whorls with 9 ribs, third with 8-10, fourth with 9 or 10 and last whorl with 6 or 7 ribs. Spiral sculpture of low, strong, narrow, nodose, primary, secondary and tertiary cords. First whorl with visible IP, P1, P2, starting s1; second whorl with SP; IP, P1, s1, P2, s2; third and fourth whorls with additional adis and occasionally with abis and tertiary cords. Last whorl with SP, adis, IP, abis, P1, s1, P2, s2, P3, P4, P5, P6, s6, ADP, MP and ABP and few tertiary cords (Fig. 10E). Primary and secondary cords of approximately similar size. Aperture moderately small, narrow, ovate. Columellar lip moderately broad with weak, low parietal tooth at adapical extremity. Rim adherent or partially erect on a small portion abapically. Anal notch deep, broad. Outer lip erect, smooth, with 6 strong, elongate denticles within, decreasing in strength abapically, consisting of ID, D1-D5. Siphonal canal very short, 12-14% of total shell length, strongly dorsally bent, broadly open. Operculum and radula unknown. Figure 14. Distribution of the species in the Cytharomorula grayi group. Cytharomorula grayi ( western Atlantic Ocean; eastern Atlantic Ocean; western Indian Ocean); C. elegantula n. sp.; C. fatuhivaensis n. sp. Figure 15 (scale bar: 500 µm) A-E. Cytharomorula arta n. sp. C. pinguis Houart, 1995)] Cytharomorula springsteeni -Tsuchiya, 2017: 954, pl. 247, fig. 4 [not C. springsteeni Houart, 1995)]. BIOCAL, stn DW66, 03/09/1985, 24°56 S, 168°22 E, 505-515 m, paratype MNHN-IM-2010-23359, 16.3 mm; M. New Caledonia, N.O. "Jean Charcot" BIOCAL, stn DW66, 03/09/1985, 24°56 S, 168°22 E, 505-515 m, paratype MNHN-IM-2010-23359, 16.3 mm; N. Protoconch, New Caledonia, Kaimon Maru Bank, SMIB 8, stn DW159, 28/01/1993, 24°46' S, 168°08' E, 241-245 m, paratype MNHN-IM-2000-34209.

  At present, 32 Recent species and one subspecies are assigned to Orania in WoRMS (MolluscaBase 2018). Of these, O. ornamentata Houart, 1995 is here transferred to Cytharomorula, and O. albozonata[START_REF] Smith | Report on the marine fauna of the Island of St. Helena[END_REF] is removed from Muricidae to the Buccinoidea (Pisaniidae).

	One of the syntypes of Cantharus albozonatus E.A.
	Smith, 1890 in the NHMUK, most probably the shell
	figured by Smith (1890: 260, pl. 21 fig. 9), is here
	illustrated (Fig. 29I-J). That species is not a muricid
	but rather a buccinid close to Enginella leucozona
	(Philippi, 1844) (K. Fraussen, in litt.). However, based
	on the images we received from A. Salvador at the
	NHMUK, other buccinids are included in the 35
	syntypes of C. albozonatus (see Fig. 29K-L), which is
	here tentatively assigned to Enginella Monterosato,
	1917.
	One new species of Orania is described herein, while
	Muricopsis carnicolor Bozzetti, 2009 (Figs 6E-F;
	21D-H) and Morula taiwana are transferred to
	Orania. This leaves 33 species and one subspecies in
	this genus (Table
	Figure 18 (scale bars: 500 µm)
	A-F. Cytharomorula fatuhivaensis n. sp., French Polynesia, Marquesas Islands, Fatu Hiva, 03/09/1990, 10°31' S,
	138°39' W, 210 m.
	A-B. Holotype MNHN-IM-2012-18829, 14.7 mm; C. Protoconch, paratype MNHN-IM-2000-34210; D.
	Paratype MNHN-IM-2000-34210, 14.8 mm; E-F. Paratype MNHN-IM-2000-34210, 15.5 mm.
	G-L. Cytharomorula manusuduirauti n. sp.
	G-H. Philippines, Aliguay Island, near 8°44' N, 123°12'40' E, sand, sandy gravel & sandy mud bottoms, 50-150
	m, holotype HMNS 2017.2223 Po.406866, 11.6 mm, photo Gary Kidder, The Houston Museum of Natural
	Science, Houston, Texas, U.S.A.; I. Philippines, Balicasag Island, near 9°31' N, 120°41' E, tangle or lumun-
	lumun net, 100-150 m, paratype HMNS 2017.2223 Po.524523, 10.1 mm, photo Gary Kidder, The Houston
	Museum of Natural Science, Houston, Texas, U.S.A.; J-L. Philippines, Aliguay Island, rocky mud, 160 m,
	paratype RH, 10.2 mm.
	M-N. Cytharomorula pinguis Houart, 1995, New Caledonia, Loyalty Ridge, 20°41'S 167°07'E, 360 m, holotype

. Shell with high spire, elongate, ovate, nodose, not or rarely exceeding 25 mm in length at maturity. Axial sculpture of last teleoconch whorl consisting of 7-9 narrow, low, sharp or weakly rounded ribs. Spiral sculpture of low, narrow, primary, secondary and tertiary cords. Aperture ovate, columellar lip weakly concave, smooth or with weak folds abapically, rim almost entirely adherent to shell. Anal notch broad, deep. Outer lip with strong, elongate denticles within. Siphonal canal short, broadly open ventrally. Radula of the type species with three dimensional rachidian tooth bearing a long, moderately broad, projecting central cusp, a small, narrow lateral denticle, a moderately long, broad, lateral cusp and strong marginal folds giving rise to small, short marginal denticles, and a bifid marginal cusp. Lateral teeth sickle shaped, with broad base. MNHN-IM-2000-952, 16.7 mm, photo Manuel Caballer (MNHN) E-Recolnat Project: ANR-11-INBS-0004.

Remarks.

  fig. 371.

	Nucella castanea (Küster, 1886) -Houart et al.,
	2010: 197.
	Type material. Purpura castanea: "There is no
	indication. The whereabouts of this type (even if from
	Kuster's coll.) are unknown. P. castanea was founded
	on material coming from F. Krauss, probably
	destroyed during the war in Stuttgart" (R. Janssen, in
	litt.); Cominella fasciata: three syntypes NHMUK
	86.4.2.10; Cominella unifasciata var. nigronodulosa:
	one syntype Oxford Museum.
	Type localities. Purpura castanea: Cape Agulhas;
	Cominella fasciata: Port Elizabeth; Cominella
	unifasciata var. concolor: Natal; Cominella
	unifasciata nigronodulosa: Port Alfred.
	Distribution. South Africa, from False Bay to the
	south coast of Natal, and Walters Shoals, south of
	Madagascar, approximately 33°12'S, 43°50'E (new
	locality).
	Other material examined. South Africa: W Cape
	Province, Witsand, 2 lv (RH); East London, 3 lv (RH)
	(radula illustrated); East Cape Province, WSW of East
	London, 6 lv (RH); Gonubie, East London, 1 lv (RH);
	Haga Haga, 3 lv & dd (RH); South Africa (no other
	data), 10 dd (RH).
	Walters Shoal seamount: MD208: stn WS08,
	33°14'S, 43°56'E, 30-33 m, MNHN-IM-2013-66431, 1
	lv (BOLD MUBA784-18; GenBank MK216551); stn
	WS08, 33°14'S, 43°56'E, 30-33 m, MNHN-IM-2013-
	66432, (BOLD MUBA785-18; GenBank MK216550)
	1 lv; stn WB09, 33°14'S, 43°56'E, 27-30 m, MNHN-
	IM-2013-66434, (BOLD MUBA786-18; GenBank
	MK216541), 1 lv; stn WB10, 33°09'S, 43°52'E, 30 m,
	MNHN-IM-2013-66424, (BOLD MUBA783-18;
	GenBank MK216555), 1 lv; stn WB10, 33°09'S,
	43°52'E, 30 m, 2 lv.

  Orania pachyrhaphe differs from B. birileffi in reaching a larger size relative to the number of teleoconch whorls; in having a lower spire; in having a spiral sculpture consisting of narrow primary cords together with weakly narrower secondary cords and small tertiary cords versus rounded, more crowded primary and secondary cords of approximately similar size; and in that adults specimens of B. birileffi lack tertiary cords. In addition, the siphonal canal of O. pachyrhaphe is relatively broader and the shell is more squamous. Its inclusion in Orania is based only on its shell morphology, which is close to O. fusulus, the type species of Orania.

	West Pacific); P. muricata (Reeve, 1846) (Indo-West
	Pacific); P. ochrostoma (Blainville, 1832) (Indo-West
	Pacific); P. ozenneana (Crosse, 1861) (Indo-West
	Pacific); P. philpoppei Houart, 2018 (Philippines and
	Japan); P. rufonotata (Carpenter, 1864) (Baja
	California, Mexico and Galapagos Islands); and P.
	submissus (E. A. Smith, 1903) (Indo-West Pacific).
	Genus Pascula Dall, 1918
	Type species by original designation: Trophon citricus
	Dall, 1908, Easter Island.
	Evokesia Radwin & D'Attilio, 1972
	Type species by original designation: Sistrum
	rufonotatum Carpenter, 1864.
	Diagnosis. Shell with high spire, broadly elongate,
	ovate, nodose, not or rarely exceeding 20 mm in
	length at maturity. Axial sculpture of last teleoconch
	whorls consisting of moderately broad, high, rounded
	ribs. Spiral sculpture of moderately high and broad
	primary cords, narrow secondary cords and few
	threads.
	Aperture ovate. Columellar lip weakly concave,
	smooth or with weak folds abapically, rim weakly
	erect abapically, adherent at small portion adapically.
	Outer lip with narrow, elongate denticles within.
	Radula of the type species illustrated by Rehder
	[1980: 135, pl. 3 (3-4)] with three dimensional
	rachidian tooth bearing a long, narrow, projecting
	central cusp, a very small, narrow lateral denticle, a
	moderately long, moderately broad, lateral cusp and
	very weak marginal folds. No marginal cusp. Lateral
	teeth sickle shaped, with broad base.
	Remarks. Eight Recent species are assigned to
	Figure 21 (scale bar: 500 µm) Pascula in WoRMS (MolluscaBase 2018): Pascula
	citrica (Dall, 1908) (Easter Island and French
	A-C. Orania fusulus (Brocchi, 1814), A. Spain, Fuengirola, RH, 22.6 mm; B. Italy, Piacenza, Piacenziano, Polynesia); P. darrosensis (E.A. Smith, 1884) (Indo-
	Castell'Arquato, Pliocene, RH, 23.2 mm; C. West Africa, Mauritania, RH, 17.4 mm.
	D-H. Orania carnicolor (Bozzetti, 2009)
	D-E. Madagascar, Lavanono, holotype MNHN-IM-2000-22900, 9.15 mm, photo Manuel Caballer (MNHN) E-
	Recolnat Project: ANR-11-INBS-0004; F-G. ATIMO VATAE, stn BP22, Madagascar secteur Ouest de
	Lavanono, 25°23.4'S, 44°51.7'E, 20-22 m, MNHN, 11.3 mm; H. Protoconch, ATIMO VATAE, stn CP3545,
	25°29'S, 46°42'E, 108-110 m, MNHN.
	I-K. Orania castanea (Küster, 1858)
	I-J. South Africa, Haga Haga, RH, 13.5 mm; K. MD208, stn WB10, Walters Shoal, 33°09,1'S, 43°51,8'E, 30 m,
	MNHN-IM-2013-66424, (BOLD MUBA783-18; GenBank MK216555), 12.4 mm.
	L. Orania gaskelli (Melvill, 1891), Papua New Guinea, Hansa Bay, RH, 16.6 mm.
	M-N. Orania serotina (A. Adams, 1853), Philippines, Cebu, Mactan Id, Punta Engaño, RH, 14.3 mm.

O-P. Usilla avenacea (Lesson, 1842), Marquesas, Ua Huka, RH, 11.1 mm. Q. Orania bimucronata

[START_REF] Reeve | Conchologia iconica , or illustration of the shells of molluscous animals[END_REF]

, Malaya, Tioman, RH, 14.6 mm.

  , P. palmeri was assigned to Morula. Although molecular genetic analyses are not yet available, the shell and radula morphology of P. palmeri (Figs 7G-H; 22O-P) is very close to P. citrica (Fig.22K-L), the type species of Pascula. Thus, while waiting for a more complete genetic analysis of this genus, we suggest assigning P. palmeri to Pascula rather than to Morula, as the type species, Morula uva (Röding, 1798), is quite different from P. palmeri.

	Genus Tenguella Arakawa, 1965
	Type species by original designation: Purpura
	granulata Duclos, 1832, Indo-West Pacific.
	Diagnosis. Shell with high spire, broadly ovate,
	nodose, rarely exceeding 30 mm in length at maturity.
	Axial sculpture of last teleoconch whorl consisting of
	8-10 broad, rounded, nodose ribs. Spiral sculpture of
	strong, high, broad primary cords and numerous
	threads, forming relatively high nodes at intersection
	with axial ribs.
	Aperture ovate. Columellar lip sinuous, smooth or
	with 1 or 2 narrow knobs abapically, rim very weakly
	erect abapically, otherwise adherent. Outer lip with
	strong denticles within.

.

  Tenguella granulata, T. chinoi n. sp. and T. ericius n. sp. are syntopic in PAPUA NIUGINI, stn PM41 and PR214, and in SANTO 2006, stn FM36; T. chinoi n. sp. and T. ericius are also syntopic in Guam (see under T. chinoi n. sp.). Tenguella granulata and T. chinoi n. sp. are syntopic in PAPUA NIUGINI stn PFM03, PM43, PR76 and PR204, and in SANTO stn VM12 and RAP15. Claremontiella consanguinea (E.A. Smith, 1891) M-N. St. Helena, lectotype NHMUK 1889.10.1.2368, 17.4 mm (photo Harry Taylor, NHMUK Photographic Unit, © Natural History Museum of London); O-P. Sao Tome, Praia Emilia, RH, 13.8 mm.Tenguella ericius n. sp. was also collected in the China Sea prior to 1960, where it is probably sympatric with T. granulata and T. chinoi n. sp., but specific locality data is missing (MNHN). As for T. chinoi n. sp., the specimens of T. ericius n. sp. collected in the China Sea are larger, reaching a length of 17.5 mm. Another shell, misidentified as M. granulata, was illustrated by[START_REF] Wells | Christmas shells. The marine molluscs of Christmas Island (Indian Ocean)[END_REF]. It is undoubtedly T. ericius. The localities given (Christmas Island and Cocos/Keelings Islands, Indian Ocean) are not unexpected, but the stated size of the shell (2 cm) is far larger than the other known specimens.

	Distribution. Eastern and western South Atlantic,
	Eastern Pacific, Gulf of California and Baja
	California, Mexico.
	Remarks.	Houart	(1997:	64)	considered
	Claremontiella consanguinea (E. A. Smith, 1891)
	(Figs 8E-F; 27M-P) a valid species and assigned it to
	Morula, together with C. nodulosa. The shell of C.
	consanguinea was separated from C. nodulosa by
	having a narrower shell with lower nodes, more
	broadly spaced spiral cords, a relatively longer
	siphonal canal and a less denticulate or smooth
	aperture.
	Figure 27 (scale bar: 500 µm) A-B. Azumamorula mutica (Lamarck, 1816), Reunion Id, harbour, 2-3 m, RH, 18.2 mm. C-L. Claremontiella nodulosa (C.B. Adams, 1845) C-D. Jamaica, lectotype MCZ 177045, 14.9 mm (photo Jennifer Trimble, curatorial assistant, MCZ); E-G. Syntypes of Ricinula ferruginosa Reeve, 1846, "W. Indies added to board", H. Cuming collection, NHMUK 1968462. E-F. 18.9 mm; G. 17.3 mm (photo Harry Taylor, NHMUK Photographic Unit, © Natural History Museum of London); H. Cape Verde, Sal Id, RH, 14.6 mm; I. Tobago, Rocky Point, Mt Irvine Bay, RH, 15.4 mm; J. Protoconch, Cuba, RH; K-L. West Africa, Angola, Cape Esterias, RH, 15.2 mm. Figs 8D-F; 11G; 27C-P; 28A-E Type species: Purpura nodulosa C.B. Adams, 1845: 2, eastern and western Atlantic (Fig. 27C-L). Other included species. Claremontiella adiakritos n. sp. and C. consanguinea (E.A. Smith, 1891) (new combination). Description. Shell moderately broad with high spire and 7 or 8 rounded, nodose axial ribs, crossed by a subsutural spiral cord and 5 primary spiral cords on convex part of teleoconch whorl, with additional, low, narrow, secondary and tertiary cords. Protoconch conical with 3.5 whorls and sinusigeral terminal notch. Aperture narrow, ovate. Columellar lip smooth or with low denticles abapically and low parietal tooth at adapical extremity, weakly concave, rim adherent to shell. Anal notch broad, moderately deep. Outer apertural lip smooth with low denticles within. Siphonal canal short, 12-14% of total shell length, narrow, open. Radula of Claremontiella of three-dimensional type consisting of a rachidian bearing a narrow, long, central cusp, a narrow, very short, lateral denticle on each side, a broad, short lateral cusp, a few marginal folds and a short marginal cusp. Lateral teeth sickle shaped, narrow with a slightly broader base (Fig. 8D-M-P. Claremontiella new genus F).

  Murichorda rumphiusi(Houart, 1996). J. Ambon, Hito, east side of Laha, paratype RH, 19.7 mm; K-L. Singapore, east coast of Park Beach, intertidally, muddy rocks, RH, 19.6 mm.

	Figure 28 A-E. Claremontiella adiakritos n. sp. A-D. West Mexico, Mazatlan. A-B. Holotype NHMUK 20180545, 23.3 mm; C-D. Paratype RH, 24.7 mm; E. West Mexico, Baja California, near San Felipe, Alecia Playa, RH, 15.2 mm. F-H. Murichorda fiscellum (Gmelin, 1791) F-G. South of west Java, Sancang, low tide, under rocks, RH, 23.2 mm; H. Aden, Gulf of Aden, RH, 26.2 mm. I. Murichorda jacobsini (Emerson & D'Attilio, 1985), Solomon Islands, Guadalcanal, Marau Sound, RH, 37.3 mm. Figs 8D; 11G; 28A-E; 30 Morula ferruginosa -Keen, 1958: 376, sp. 411; Abbott, 1974: 178, fig. 1875 (not Ricinula ferruginosa Reeve, 1846). Morula (Morunella) ferruginosa -Keen, 1971: 554, fig. 1092; Victor Alamo & Violeta Valdivieso, 1997: 54 (not Ricinula ferruginosa Reeve, 1846). Evokesia ferruginosa -Radwin & D'Attilio, 1972: 338, fig. 1A; Radwin & D'Attilio, 1976: 143, pl. 3, fig. 5; Kaicher, 1979, card 2038 (not Ricinula ferruginosa Reeve, 1846). Pascula ferruginosa -Vokes, 1984: 2014, 215 (not Ricinula ferruginosa Reeve, 1846). Morula (Morula) ferruginosa -Skoglund, 2002: 116 (not Ricinula ferruginosa Reeve, 1846). Type material. Holotype NHMUK 20180545, West Mexico, Mazatlan, lv. Paratypes: West Mexico, Mazatlan, lv, 2 RH; Mexico, Baja California Sur, Playa Santispac, about 30 km south of Mulege, 2 IRSNB I.G. 33893; MT. 3744, 2 MNHN-IM-2000-34221 . Type locality. West Mexico, Mazatlan. Other Material examined. Baja California, Mexico, Alecia Playa, near San Felipe, 24 lv & dd, RH; Guaymas, 3 lv & dd, RH (radula illustrated); Mexico, Adair Bay, 2 lv, RH; Gulf of California (no other data), 3 lv, RH; Mexico, Mulege, 2 dd, RH; Baja California, Mexico, Puertocitos, 1 dd, RH; Baja J-L. Claremontiella adiakritos n. sp. California, Mexico, La Paz, 2 lv, RH.

  3. Lanceolate, narrow, heavy, nodose. Subsutural ramp broad, strongly sloping, concave. Radula of three-dimensional type consisting of a rachidian bearing a narrow, long, central cusp, a narrow, very short, lateral denticle on each side, a broad, shorter lateral cusp and few marginal folds. Lateral teeth sickle shaped, narrow with a slightly broader base. Muricodrupa fenestrata[START_REF] Blainville | Disposition méthodique des espèces récentes et fossiles des genres Pourpre, Ricinule, Licorne et Concholepas de M. de Lamarck, et description des espèces nouvelles ou peu connues, faisant partie de la collection du Muséum d'Histoire Naturelle de Paris[END_REF], New Caledonia, Ouvea, RH, 30.3 mm. C-D. Muricodrupa anaxares[START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF], South Africa, Durban, Reunion rocks, low tide, in rock crevices, RH, 14.6 mm. E-F. Lauta parva(Reeve, 1845), Philippines, Cebu, RH, 9.4 mm. G-H. Engina alveolata[START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF], Fiji, Wayasawa group, Wasayama Id, under coral slab, low tide, RH, 12.9 mm. Remarks.[START_REF] Vokes | Comparison of the Muricidae of the Eastern Pacific and Western Atlantic, with cognate species[END_REF] figured Reeve's illustrated syntype of Ricinula ferruginosa(NHMUK 1968462) and noted that the syntypes were specimens of the Atlantic Trachypollia nodulosa (C.B. Adams), leaving the Pacific species without a name.[START_REF] Vokes | Comparison of the Muricidae of the Eastern Pacific and Western Atlantic, with cognate species[END_REF] also illustrated the lectotype of T. nodulosa. Claremontiella adiakritos n. sp. is assigned to Claremontiella because the shell and radula morphology are similar to those of C. nodulosa, type species of the genus, and to C. consanguinea. The species has a very high spire with a low subsutural spiral cord; low, narrow cords on subsutural ramp; five low, broad, spiral cords on convex part of teleoconch whorl; numerous spiral threads; and a very short siphonal canal. The outer apertural lip bears low denticles within, and is occasionally smooth.

	Dark brown or blackish-brown, white between nodes
	of first primary cord (P1), also on other cords but less
	obvious. Aperture bluish-white within with brown
	band within outer lip up to white apertural denticles.
	Columellar lip bluish-white with a darker area
	abapically.
	Spire very high with teleoconch up to 6 weakly
	convex, narrow, angulate, shouldered, nodose whorls.
	Suture adpressed. Protoconch in examined material
	unknown, eroded in all specimens.
	Axial sculpture of teleoconch whorls consisting of
	low, weak, narrow, rounded ribs with moderately
	high, sharp nodes at intersection of primary spiral
	cords. First and second whorls eroded in all
	specimens, third whorl with 10 ribs, fourth with 9,
	fifth with 9 or 10, last whorl with 7 ribs. Spiral
	sculpture of low, rounded, broad, nodose primary
	cords and numerous threads. Last teleoconch whorl
	with SP, P1-P5 and additional threads between each
	pair of cords. P1 broadest, P2-P3 weakly narrower,
	similar in strength, P4 narrower and lower, P5
	smallest on adapical part of siphonal canal.
	Aperture small, ovate. Columellar lip narrow, smooth,
	rim adherent, with very weak low parietal tooth at
	adapical extremity. Anal notch shallow, broad. Outer
	lip smooth with low, small denticles within, consisting
	of D1-D4, strongly decreasing in strength abapically;
	very low ID occasionally observed. Denticles
	sometimes obsolete. Siphonal canal very short, 10-
	12% of total shell length, narrow, weakly dorsally
	recurved, broadly open.
	Operculum dark brown, ovate with subapical nucleus
	in lower right.
	Figure 29
	A-B.

I-L. Cantharus (Tritonidea) albozonatus Smith, 1890, St. Helena, W.H. Turton collection. I-J.

Syntype NHMUK 1889.10.1.2356-2361, 15.8 mm; K-L. Syntype NHMUK 1889.10.1.152-161, 10.9 mm.

  reported this species (without any figure) from Bocapán and El Rubio, Tumbes, Peru, but Claremontiella adiakritos n. sp. has been frequently confused with Trachypollia lugubris and no voucher material was referenced (V. Mogollon, in litt.). Therefore, the presence of C. adiakritos in Peru and the Galapagos Islands remains doubtful. Claremontiella nodulosa differs from C. adiakritos n. sp. in having a shell with slightly larger primary spiral cords with rounded nodes instead of more sharp knobs, and in having a generally less high spire, a broader, obvious and nodose subsutural cord, and a darker coloured aperture with thicker outer apertural lip and more obvious denticles within.

Etymology. Adiakritos (G) meaning mixed, undistinguishable, named for its resemblance to some forms of Claremontiella nodulosa (C.B.

[START_REF] Adams | Speciorum novarum conchyliorum Jamaica repertorum synopsis[END_REF]

, with which it has often been confused in literature.

  Shell large, up to 44.5 mm in length, broadly ovate, weakly angulate, shouldered, with 6-8 broad, high, rounded axial ribs, crossed by 5 high, longitudinally grooved primary spiral cords, occasionally deeply excavated between each pair of cords. Intersection of axial ribs and primary spiral cords forming low, broad knobs. Aperture broadly ovate with low denticles within outer lip. Siphonal canal short to moderate in length, 13-29% of total shell length, broadly open. Radula ergalataxine (Figs 8G-H; 9A), of threedimensional type, consisting of a narrow, long central cusp, a very short, narrow, lateral denticle on each side, a broad, moderately long lateral cusp and occasionally a few low marginal folds. Lateral teeth sickle shaped, broad.

(new combination) and Murichorda jacobsini (Emerson & D'Attilio, 1985) (new combination).

Description.

  Muri, from Muricidae + suffix chorda (L): rope, twine. Named for the strong, broad spiral cords, a characteristic shared by the shells assigned to this new genus. Ricinula parvaReeve, 1846: pl. 6, fig. 43. Description. Shell small, up to 10 mm in length, weakly ovate and shouldered, nodose, with 7 or 8 axial ribs crossed by narrow, rounded, primary cords consisting of SP near suture, P1-P5 on convex part of teleoconch whorl and ADP on siphonal canal, with rounded, high nodes at intersection of axial ribs and spiral cords. Spire high with conical protoconch of 3.5 whorls. Aperture strongly ovate, narrow, with strong denticles within outer lip. Columellar lip narrow, almost straight, smooth, rim almost entirely adherent to shell. Anal notch broad, moderately deep. Siphonal canal short, 10-13% of total shell length. Radula unknown. Lauta parva is sister to all other members of subclade Z in Claremont et al (2013: fig. 1-3B), suggesting that a new genus is required for this species. The genus Morula Schumacher, 1817 in which Lauta parva was previously included consisted of 19 species. One of them, M. anaxares, is now assigned to Muricodrupa Iredale, 1918; two species, M. nodulosa and C. consanguinea, are now assigned to Claremontiella n. gen.; one species, M. rumphiusi, is now assigned to Murichorda n. gen.; and one species, M. parva, is here assigned to Lauta n. gen. Of the 14 remaining species of Morula s.s., six were analysed by Claremont et al (2013): M. aspera (Lamarck, 1816), M. chrysostoma (Deshayes, 1844), M. echinata (Reeve, 1846) (= Sistrum ventricosulum G. & H. Nevill, 1875, new synonymy) (as M. benedicta in Claremont et al., 2013), M. nodicostata (Pease, 1868), M. uva (Röding, 1798) and M. zebrina Houart, 2004. The eight remaining species are here provisionally retained in Morula s.s. (Table 2), pending future genetic analyses. Etymology. The generic name Lauta (L: neat, elegant) is derived from Ricinula lauta Reeve, 1846, junior synonym of Engina alveolata[START_REF] Kiener | Spécies général et iconographie des coquilles vivantes[END_REF] (Fig.29G-H), a species of Pisaniidae (Buccinoidea) whose shell morphology and (particularly) colouration is strangely reminiscent of the type species, L. parva.

	Lauta new genus Fig. 29E-F Type species *Lataxiena Jousseaume, 1883 Genus Lataxiena lataxiena Jousseaume, 1883 = Trophon fimbriatus Hinds, 1844 *Lauta n. gen. Ricinula parva Reeve, 1846 *L. parva (Reeve, 1846) Species L. blosvillei (Deshayes, 1832) L. bombayana (Melvill, 1893) L. cumella (Jousseaume, 1898) L. desserti Houart, 1995 L. habropenos Houart, 1995 L. lutescena Zhang & Zhang, 2015 L. solenosteiroides Houart, Fraussen & Barbier, 2013 Lindapterys Petuch, 1987 Lindapterys vokesae Petuch, 1987 (early Miocene, Florida) L. domlamyi Garrigues & Merle, 2014 L. murex (Hedley, 1922) L. sanderi Petuch, 1987 L. soderiae Callea, Volpi, Martignoni & Borri, 2001 Table 2 (continued) Genus Type species Species *Orania Pallary, 1900 Note: Type species not analysed in Claremont et al. (2013) Murex spadae Libassi, 1859 = Murex fusulus Brocchi, 1814 O. adiastolos Houart, 1995 O. archaea archaea Houart, 1995 O. archaea hitomiae Houart & Moe, 2011 O. atea Houart & Tröndlé, 2008 Type species: Table 2 (continued) *L. fimbriata (Hinds,1844) O. badia	NC NC NC

Distribution. Philippine Islands.

Remarks.

ACKNOWLEDGEMENTS

Most of the material in this paper originates from numerous shore-based expeditions and deep water cruises, conducted between 1977 and 2017 by the MNHN and Pro-Natura International (PNI) as part of the Our Planet Reviewed programme (SANTO 2006, ATIMO VATAE, MIRIKY, MAINBAZA, PAPUA NIUGINI, PI Philippe Bouchet; KANACONO, PI Nicolas Puillandre; KANADEEP, PI Sarah Samadi), and/or by MNHN and Institut de Recherche pour le Développement (IRD) as part of the Tropical Deep-Sea Benthos programme in New Caledonia (BIOCAL, CHALCAL 2, SMIB 3, 4, 5, 8, BERYX 11, BATHUS 2, NORFOLK 1, NORFOLK 2, EBISCO, TERRASSES, EXBODI), Wallis & Futuna (MUSORSTOM 7), Vanuatu (MUSORSTOM 8), the Marquesas (MUSORSTOM 9), Tonga (BORDAU 2), the Philippines (PANGLAO 2005, AURORA 2007), the Mozambique Channel (BIOMAGLO) and Walters Shoal (PIs Claude Lévi, Bertrand Richer de Forges, Patrick Lehodey, Cécile Debitus, Philippe Bouchet, Sarah Samadi, Laure Corbari). Additional sources are BENTHEDI (PI Bernard Thomassin), the Marion-Dufresne MD32 cruise off Réunion (PI Alain Guille), the LIFOU 2000 expedition to the Loyalty Islands, the PANGLAO 2004 expedition to the Philippines, and the PAKAIHI I TE MOANA expedition to the Marquesas Is. Scientific partners included the University of Papua New Guinea (UPNG), University San Carlos (Cebu City) and the Bureau of Fisheries and Aquatic Resources (BFAR, Manila), Institut d'Halieutique et des Sciences Marines, University of Toliara (IH.SM) and the Madagascar bureau of Wildlife Conservation Society (WCS), Instituto Español de Oceanografia, the Marine and Polar Programme of the International Union for Conservation of Nature (IUCN International), and Agence des Aires Marines Protégées (AAMP), with access to ship time through the Flotte Océanographique Française, and funding from the Total Foundation, Prince Albert II of Monaco Foundation, Stavros Niarchos Foundation, Vinci Entrepose Contracting, Fondation EDF, the French Global Environment Fund (FFEM), Fonds Pacifique and the Government of New Caledonia. For station lists and context of the expeditions, see https://expeditions.mnhn.fr/ and Bouchet et al. (2008). We are very grateful to Philippe Bouchet (Muséum national d'Histoire naturelle, Paris, France) for the opportunities he has given us to study the material collected during the Tropical Deep Sea Benthos campaigns in the Indo-West Pacific, for information about the various expeditions and for constructive comments. We are also very thankful to Barbara Buge, Virginie Héros and Philippe Maestrati (MNHN) for their help in searching for and sending specimens in loan, and for relevant information. Manuel Caballer (MNHN) provided the images of the MNHN holotypes, E-Recolnat Project: ANR-11-INBS-0004. Thanks also to everyone who contributed to this work by sending photos and loans, and for other information. Without their help, it would have been difficult to complete this article. Our sincerest thanks to Adam Baldinger and Marcia Kazmierczak (Museum of Comparative Zoology, Harvard University, Cambridge, U.S.A) for the image of the lectotype of Purpura nodulosa C.B. Adams, 1845; to Andrea Barco, GEOMAR, Helmholtz Centre for Ocean Research, Kiel, Germany for information about genetic analyses; to Philippe Bouchet (MNHN), Anders Warén (Natural History Museum, Stockholm, Sweden) and Yuri Kantor (A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia) for preparation of radulae and for SEM work. to Martine Claremont (Boston, Massachusetts, U.S.A) for reading, commenting on and correcting the first draft of this ms; to Jacques Colomb (Marseille, France) for providing me with specimens of C. nodulosa from Virgin Islands and C. consanguinea from São Tomé for genetic analysis; to Dai Herbert, Linda Davis and the late Richard (Dick) Kilburn (KwaZulu-Natal Museum, South Africa) for the loan of material; to Roland Janssen (Forschungsinstitut Senckenberg, Frankfurt, Germany) for information about the type material of Purpura castanea Küster, 1858; to Bruce Marshall (Museum of New Zealand Te Papa Tongarewa, Wellington, New Zealand) for sending us soft parts of Pascula palmeri for study and SEM of the radula; to Shawn Miller (Okinawa, Japan) for providing me with some specimens of Orania pachyrhaphe a few years ago; to Valentin Mogollón Avila (Universidad Nacional Federico Villarreal, Lima, Peru) for information about the possible presence of Claremontiella adiakritos n. sp. in Peru; to Tina Petway and Gary Kidder (The Houston Museum of Natural Science, Houston, Texas, U.S.A.) for information and photos of the holotype and paratypes of Cytharomorula manusuduirauti n. sp.; to Jane Pickering (then assistant curator, zoological collections at the Oxford University Museum, United Kingdom), for the loan of the syntype of Cominella unifasciata nigronodulosa Turton, 1932Turton, in 1995;; to Andreia Salvador and Harry Taylor (Natural History Museum, United Kingdom), for providing information and photographs of the type material deposited in NHMUK; and to Emmanuel Tardy (Muséum d'histoire naturelle, Genève, Switzerland) for information about a type from Kiener.

Chalut de loutre Drague Charcot (Charcot dredge) DW: Drague Warén (Warén dredge) PM: M = marées (tides) PR: R = récoltes à vue (handpicking) Specimens dd: empty shell(s). lv: live collected specimen(s) *Bedevina Habe, 1946

Table 2. List of the valid extant taxa in ERGALATAXINAE Bold: new taxa described in this paper. Asterisk: taxa sequenced in [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF] or in this paper. Question mark: doubtful classification after sequencing, but a temporary placement in the genera in which they are currently assigned is suggested. NC: New combination.

Genus

Type species Species

Azumamorula Emerson, 1968 Ricinula mutica Lamarck, 1816

A. mutica (Lamarck, 1816) U. ahiparana (Powell, 1927) U. marwicki Powell, 1952U. williamsi Powell, 1952 ________________________ Note: Small genetic distances are suggestive of possible synonymy of species; for example, there was < 1% divergence among COI sequences for the pair Morula uva and M. aspera, and similarly for the pair Cronia amygdala and C. aurantiaca [START_REF] Claremont | A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae)[END_REF].