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Hani Hanjour

Hani Salih Hasan Hanjour (هاني صالح حسن حنجور; 30 August 1972 – 11 September 2001) was a Saudi Arabian terrorist who was the hijacker-pilot of American Airlines Flight 77, crashing the plane into the Pentagon as part of the 9/11 attacks in 2001.

Hanjour first went to the United States in 1991, enrolling at the University of Arizona, where he studied English for a few months before returning to Saudi Arabia early the next year. He returned to the United States in 1996, studying English in California before he began taking flying lessons in Florida and then Arizona. He received his commercial pilot certificate in 1999, and went back to his native Saudi Arabia to find a job as a commercial pilot. Hanjour applied to civil aviation school in Jeddah, but was turned down. Hanjour left his family in late 1999, telling them that he would be traveling to the United Arab Emirates to find work. According to Khalid Sheikh Mohammed, Osama bin Laden or Mohammed Atef identified Hanjour at an Afghanistan training camp as a trained pilot and selected him to participate in the 11 September attacks.

Hanjour arrived in the United States again in December 2000. He joined up with Nawaf al-Hazmi in San Diego. They immediately left for Arizona, where Hanjour engaged in refresher pilot training. In April 2001, they relocated to Falls Church, Virginia and then Paterson, New Jersey in late May where Hanjour took additional flight training.

Hanjour returned to the Washington, D.C., metropolitan area on 2 September 2001, checking into a motel in Laurel, Maryland. On 11 September, Hanjour boarded American Airlines Flight 77, took control of the aircraft after his team of hijackers helped subdue the pilots, passengers, and crew, and flew the plane into the Pentagon as part of the 11 September attacks. The crash killed all 64 passengers on board the aircraft and 125 people in the Pentagon.

While in Florida and Arizona, Hanjour befriended and trained with Rayed Mohammed Abdullah Ali, a fellow Saudi Arabian who emigrated to Manawatū-Whanganui in 2006 to train as a pilot. He was deported from his Palmerston North home after his links to Hanjour were exposed.

Early life and education
Hanjour was one of seven children, born to a food-supply businessman in Ta'if, Saudi Arabia. During his youth, Hanjour wanted to drop out of school to become a flight attendant, although his brother Abdulrahman discouraged this route, and tried to help him focus on his studies.

According to his eldest brother, Hanjour traveled to Afghanistan in the late 1980s as a teenager to participate in the conflict against the Soviet Union. The Soviets had already withdrawn by the time he arrived in the country and he instead worked for a relief agency.

Early 1990s
Hanjour was the first to arrive in the United States, much earlier than other hijackers. He first came to the United States in 1991 to study English at the University of Arizona's Center for English as a Second Language. Hanjour's eldest brother Abdulrahman helped him apply to the eight-week program, and found a room in Tucson, Arizona, for Hanjour near the Islamic Center of Tucson. Hanjour arrived for the English language program on 3 October 1991, and stayed until early February 1992, when he returned to Saudi Arabia. Hanjour shared a three-bedroom home on the corner of 4th Avenue and 4th Street owned and managed by a father-son team, who made a living renovating and renting rooms to international students and devoting their energies to spreading a born-again Christian influence; Bob, the oldest son, lived in this house and rented the room directly to Hanjour. Hanjour was a model housemate; he was extremely respectful of others, apolitical in his points of view, and appeared as a nonchalant happy-go-lucky teenager with very weak English-speaking skills. Hanjour claimed that he was interested in being an airplane mechanic and claimed that such a position was considered highly in Saudi Arabia. Hanjour participated in morning, noon, and evening prayers at the local mosque. As early as Hanjour moved into this house, he was under the constant watch of two "uncles" who would pick him up for the weekends so that he would spend time with them within their circle, thereby minimizing his contact with his American housemates and friends from the university. In December 1991 Hanjour informed Bob that he missed Saudi Arabia and would be leaving the United States due to homesickness. Hanjour was the only hijacker to visit the United States prior to any intentions for a large-scale attack and was not linked to the Hamburg cell in Germany, which composed of the three hijackers who were taking flight lessons to become pilots. He was the first to receive a license out of the three other men, most likely due to arriving to the U.S. prior to the preparations for the attacks.

Over the next five years, Hanjour remained in Saudi Arabia, helping the family manage a lemon and date farm near Ta'if. His family often reminded Hanjour that he was getting past the age where he ought to get married and start a family, but Hanjour insisted he wanted to settle down more. While in Saudi Arabia, Hanjour applied for a job with Saudi Arabian Airlines, but was turned down due to poor grades. The airline told Hanjour they would consider him if he obtained a commercial pilot's license in the United States.

1996
In April 1996, Hanjour returned to the United States, staying with family friends, Susan and Adnan Khalil, in Miramar, Florida, for a month before heading to Oakland, California, to study English and attend flight school. Hanjour was admitted to the Sierra Academy of Aeronautics, but before beginning flight training, the academy arranged for Hanjour to take intensive English courses at ESL Language Center in Oakland. The flight school also arranged for Hanjour to stay with a host family, with whom he moved in on 20 May 1996. Hanjour completed the English program in August, and in early September 1996, he attended a single day of ground school courses at the Sierra Academy of Aeronautics before withdrawing, citing financial worries about the $35,000 cost.

Hanjour left Oakland in September and moved to Phoenix, Arizona, paying $4,800 for lessons at CRM Flight Cockpit Resource Management in Scottsdale. Receiving poor marks, Hanjour dropped out of flight school, and returned to Saudi Arabia at the end of November 1996.

Late 1990s
Hanjour re-entered the United States on 15 November 1997, taking additional English courses in Florida, then returning to Phoenix, where he shared an apartment with Bandar al-Hazmi.

After arriving in Florida on November 1997, Hanjour met a man Rayed Mohammed Abdullah Ali, a mutual friend through Bandar al-Hazmi. Al-Hamzi had suggested that Ali train to be a pilot in Florida. Hanjour and Ali subsequently trained together both in Florida and Arizona, and became friends. Ali was a leader at the Islamic Cultural Centre in Phoenix where, the FBI says, he "reportedly gave extremist speeches at the mosque". This, however, was disputed by a mosque staff member, who told the New Zealand Herald in June 2006 that Ali "was never a leader for the mosque and he never gave speeches at the mosque".

The 9/11 Commission Report reported that Abdullah attended the same Phoenix flight school as Hanjour and records show the pair used a flight simulator together on 23 June 2001. Ali later emigrated to New Zealand, where he settled in Auckland and then in Palmerston North to train as a pilot. He was identified as a friend and colleague of Hanjour in July 2006, less than six months after he first arrived in New Zealand. Immigration Minister David Cunliffe said that Ali "was directly associated with persons responsible for the terrorist attacks in the United States on Sept. 11, 2001".

In December, he resumed training at CRM Flight Cockpit Resource Management for a few weeks, before pursuing training at Arizona Aviation. Bandar al-Hazmi and Hanjour stayed in Arizona, continued taking flight lessons at Arizona Aviation throughout 1998 and early 1999. After moving out of Bandar's place in March, Hanjour lived in several apartments in Tempe, Mesa and Phoenix. In February, financial records showed that Hanjour had taken a trip to Las Vegas, Nevada. In addition to flight training at Arizona Aviation, Hanjour enrolled in flight simulator classes at the Sawyer School of Aviation where he made only three or four visits. Lotfi Raissi would begin taking lessons at the same school a month after Hanjour quit, part of what piqued the FBI's interest in Raissi.

An FBI informant named Aukai Collins claims he told the FBI about Hanjour's activities during 1998, giving them Hanjour's name and phone number, and warning them that more and more foreign-born Muslims seem to be taking flying lessons. The FBI admits it paid Collins to monitor the Islamic and Arab communities in Phoenix at the time, but denies Collins told them anything about Hanjour.

Hanjour gained his FAA commercial pilot certificate in April 1999, getting a "satisfactory" rating from the examiner. After receiving his commercial pilot certificate, Hanjour went on to train at other flight schools for more advanced flying. This caused concern among the instructors as they thought “he was so bad a pilot and spoke such poor English that they contacted the Federal Aviation Administration to verify that his license was not a fake.” The FAA verified his license and offered to provide him with English language tutors. Hanjour's bank records indicate that he travelled to Ontario, Canada, in March 1999 for an unknown reason.

He traveled to Saudi Arabia to get a job working with Saudi Arabian Airlines as a commercial pilot but was rejected by a civil aviation school in Jeddah. His brother, Yasser, relayed that Hanjour, frustrated, "turned his attention toward religious texts and cassette tapes of militant Islamic preachers." He told his family in late 1999 he was heading to the United Arab Emirates to find work. However, it is likely that he headed to al-Qaeda training camps in Afghanistan. He was known by al-Qaeda as Urwa al-Taa'ifi.

2000
In May 2000, a third person accompanied Nawaf al-Hazmi and Khalid al-Mihdhar to Sorbi's Flying Club where he waited on the ground as they took a flight lesson. It has been theorized this may have been Hanjour.

In September Hanjour again sent his $110 registration to the ELS Language Center, which leased space on Holy Names College campus in Oakland, California, to continue his English studies. He also applied for another U.S. student visa. Although he was accepted, after the attacks, it would be reported that his visa application was 'suspicious'. Granted an F-1 student visa in Jeddah, Saudi Arabia, September 2000, he failed to reveal that he had previously traveled to the U.S. He never turned up for classes at the ELS Language Center, and when the school contacted its Saudi representative, he reported that he could not find Hanjour either.

On 5 December, Hanjour opened a CitiBank account in Deira, Dubai. On 8 December, Hanjour was recorded flying into the Cincinnati/Northern Kentucky International Airport, and is thought to have met with Nawaf al-Hazmi in San Diego shortly thereafter.

2001
Hanjour came back to San Diego in December 2000, frequently visiting Abdussattar Shaikh's house, which was shared with Nawaf al-Hazmi and Khalid al-Mihdhar. During this time Hanjour may have visited the San Diego Zoo in February, as a security guard recalls having to page his name to reclaim a lost briefcase containing cash and Arabic documents and later recognized his photograph. Shortly afterwards, the three hijackers moved out of Shaikh's house to Falls Church, Virginia.

The ELS Language Center at Oakland University said Hanjour reached a level of proficiency sufficient to "survive very well in the English language". However, in January 2001, Arizona JetTech flight school managers reported him to the FAA at least five times because his English was inadequate for the commercial pilot certificate he had already obtained. It took him five hours to complete an oral exam meant to last just two hours, said Peggy Chevrette. Hanjour failed UA English classes with a 0.26 GPA and a JetTech manager said "He could not fly at all." The certificate was a requirement for him to join the Saudi Arabian pilot's academy. His FAA certificate had become invalid late in 1999 when he failed to take a mandatory medical examination. In February, Hanjour began advanced simulator training in Mesa, Arizona.

He and Hazmi moved out of Mesa at the end of March, and they were in Falls Church, Virginia, by 4 April. Falls Church was the location of the Dar al-Hijrah mosque in the Washington, D.C., metropolitan area. Anwar Al-Awlaki was the recently appointed new Imam whom Hazmi had met with in San Diego.

At the mosque, Hanjour and Hazmi soon met Eyad Alrababah, a Jordanian who later pleaded guilty to document fraud and was deported. They had told him that they were looking for an apartment to rent, and he found a friend who rented them an apartment in Alexandria where they stayed. On 4 April 2001, Hanjour asked to forward his utility deposits to 3159 Row Street, Falls Church, Virginia, which was the same address as the mosque.

When police raided the Hamburg apartment of Ramzi bin al-Shibh (the "20th hijacker") while investigating the 9/11 attacks, Awlaki's telephone number was found among bin al-Shibh's personal contact information.

On 2 May 2001, two new roommates joined them in Virginia: Majed Moqed and Ahmed al-Ghamdi, both of whom had just flown into the United States from the Middle East.

Alrababah later suggested they all go together to look at apartments in Fairfield, Connecticut. On May 8, Alrababah, Hanjour, Hazmi, Moqed and Ghamdi traveled to Fairfield to look for housing. While there, they also called several local flight schools. They then travelled briefly to Paterson to look at that area as well. Rababah has contended that, after this trip, he never saw any of the men again.

Sometime at the end of May 2001, Hanjour rented a one-bedroom apartment in Paterson, New Jersey. He lived there with at least one roommate and was visited by several other hijackers, including Mohamed Atta. During his time in New Jersey, he and Hazmi rented three different cars including a sedan in June that Hanjour cosigned with the alias "Hani Saleh Hassan". He later made his last phone call to his family back in Saudi Arabia, during which he claimed to be phoning from a payphone in the United Arab Emirates, where he was supposedly still working.

Hanjour, along with at least five other future hijackers, are believed to have traveled to Las Vegas several times in mid-2001, where they allegedly drank alcohol, gambled, and visited lap dancing clubs.

On 20 July, Hanjour flew to the Montgomery County Airpark in Maryland from New Jersey on a practice flight with fellow hijacker Nawaf Alhazmi.

On 1 August, Hanjour and Almihdhar returned to Falls Church to obtain fraudulent documentation at a 7-Eleven convenience store where an illegal side business operated for such a service. There they met Luis Martinez-Flores, himself also an illegal immigrant, who agreed to help them for a $100 fee. They drove together to a DMV office at a mall in nearby Springfield, Virginia, where Martinez-Flores gave them a false address in Falls Church to use, and signed legal forms attesting that they lived there. Hanjour and Almihdhar were then granted state identity cards. (Martinez-Flores was later sentenced to 21 months in prison for aiding them, and giving false testimony to police.) On that same day, Hanjour was stopped by police for driving a Toyota Corolla 55 mph in a 30 mph zone in Arlington, Virginia, for which he paid a $70 fine.

Employees at Advance Travel Service in Totowa, New Jersey later claimed that Moqed and Hanjour had both purchased tickets there. They claimed that Hanjour spoke very little English, and Moqed did most of the speaking. Hanjour requested a seat in the front row of the airplane. Their credit card failed to authorize, and after being told the agency did not accept personal checks, the pair left to withdraw cash. They returned shortly afterwards and paid the $1,842.25 total in cash.

Hanjour began making cross-country flights in August to test security, and tried to rent a plane from Freeway Airport in Maryland; though he was declined after exhibiting difficulty controlling and landing a single-engine Cessna 172. He moved out of his New Jersey apartment on September 1, and was photographed four days later using an ATM with fellow hijacker Majed Moqed in Laurel, Maryland, where all five Flight 77 hijackers had purchased a 1-week membership in a local Gold's Gym. There, Hanjour claimed that his first name translated as warrior when a gym employee asked if there was an English translation of their Arabic names. (Hani actually translates as "contented.")

On 10 September 2001, Hanjour, Mihdhar, and Hazmi checked into the Marriott Residence Inn in Herndon, Virginia where Saleh Ibn Abdul Rahman Hussayen, a prominent Saudi government official, was staying. No evidence was ever uncovered that they had met, or knew of each other's presence.

9/11 attacks
At 7:35 a.m. on 11 September 2001, Hanjour arrived at the passenger security checkpoint at Washington Dulles International Airport, 26 miles west of Washington, D.C., en route to board American Airlines Flight 77.

Some earlier reports stated he may not have had a ticket or appeared on any manifest, however he was documented by the 9/11 Commission as having been assigned to seat 1B in first class, and reported to have bought a single first-class ticket from Advance Travel Service in Totowa. In the security tape footage released in 2004, Hanjour appears to walk through the metal detector without setting it off, which likely means that agents at the terminal were not looking at any warning signal that indicated if he had weapons. With this, it allowed Hanjour to board Flight 77 without any authorization. The flight was scheduled to depart at 8:10, but ended up departing 10 minutes late from Gate D26 at Dulles. The last normal radio communications from the aircraft to air traffic control occurred at 08:50:51. At 08:54, Flight 77 began to deviate from its normal, assigned flight path and turned south, and then hijackers set the flight's autopilot heading for Washington, D.C. Passenger Barbara Olson called her husband, United States Solicitor General Ted Olson, and reported that the plane had been hijacked and that the assailants had box cutters and knives. Using the flight intercom, Hanjour announced the flight was hijacked. As Flight 77 was 5 miles (8.0 km) west-southwest of the Pentagon, it made a 330-degree turn. At the end of the turn, it was descending through 2,200 feet (670 m), pointed toward the Pentagon and downtown Washington. Hanjour advanced the throttles to maximum power and dove towards the Pentagon at a speed of over 530 mph. At 09:37:46, Hanjour crashed the Boeing 757 into the west façade of the Pentagon, killing himself and all 63 passengers and 6 crew members aboard along with 125 on the ground in the Pentagon. While level above the ground and seconds from the crash, the airplane's wings knocked over light poles and its right engine smashed into a power generator, creating a smoke trail seconds before smashing into the Pentagon. In the recovery process at the Pentagon, remains of all five Flight 77 hijackers were identified through a process of elimination, as not matching any DNA samples for the victims, and put into custody of the FBI.

Family denial
After the 11 September attacks, Hanjour's family in Saudi Arabia vehemently stated that they could not, and would not, believe he had been involved as one of the hijacker pilots, and also stated that he had phoned them just eight hours prior to the hijackings and his voice did not sound strange or unusual at all.

In popular culture

 * Afghan-Canadian actor Qaseem Gul portrayed Hani Hanjour in the Canadian TV series Mayday Season 16: Episode 2 (2016) called "9/11: The Pentagon Attack" and Air Crash Investigation Special Report Season 2: Episode 1 (2019) called "Headline News".

title: Hani Hanjour


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Hyena

Hyenas or hyaenas (from Ancient Greek ὕαινα, hýaina) are feliform carnivoran mammals belonging to the family Hyaenidae. With just four extant species (each in its own genus), it is the fifth-smallest family in the order Carnivora and one of the smallest in the class Mammalia. Despite their low diversity, hyenas are unique and vital components of most African ecosystems.

Although phylogenetically closer to felines and viverrids, hyenas are behaviourally and morphologically similar to canids in several elements due to convergent evolution: both hyenas and canines are non-arboreal, cursorial hunters that catch prey with their teeth rather than claws. Both eat food quickly and may store it, and their calloused feet with large, blunt, nonretractable claws are adapted for running and making sharp turns. However, hyenas' grooming, scent marking, defecation habits, mating and parental behavior are consistent with the behavior of other feliforms.

Hyenas feature prominently in the folklore and mythology of human cultures that live alongside them. Hyenas are commonly viewed as frightening and worthy of contempt. In some cultures, hyenas are thought to influence people's spirits, rob graves, and steal livestock and children. Other cultures associate them with witchcraft, using their body parts in traditional medicine.

Origins
Hyenas originated in the jungles of Miocene Eurasia 22 million years ago, when most early feliform species were still largely arboreal. The first ancestral hyenas were likely similar to the modern African civet; one of the earliest hyena species described, Plioviverrops, was a lithe, civet-like animal that inhabited Eurasia 20–22 million years ago, and is identifiable as a hyaenid by the structure of the middle ear and dentition. The lineage of Plioviverrops prospered, and gave rise to descendants with longer legs and more pointed jaws, a direction similar to that taken by canids in North America. Hyenas then diversified into two distinct types: lightly built dog-like hyenas and robust bone-crushing hyenas. Although the dog-like hyenas thrived 15 million years ago (with one taxon having colonised North America), they became extinct after a change in climate, along with the arrival of canids into Eurasia. Of the dog-like hyena lineage, only the insectivorous aardwolf survived, while the bone-crushing hyenas (including the extant spotted, brown and striped hyenas) became the undisputed top scavengers of Eurasia and Africa.

Rise and fall of the dog-like hyenas
The descendants of Plioviverrops reached their peak 15 million years ago, with more than 30 species having been identified. Unlike most modern hyena species, which are specialised bone-crushers, these dog-like hyenas were nimble-bodied, wolfish animals; one species among them was Ictitherium viverrinum, which was similar to a jackal. The dog-like hyenas were numerous; in some Miocene fossil sites, the remains of Ictitherium and other dog-like hyenas outnumber those of all other carnivores combined. The decline of the dog-like hyenas began 5–7 million years ago during a period of climate change, exacerbated by canids crossing the Bering land bridge to Eurasia. One species, Chasmaporthetes ossifragus, managed to cross the land bridge into North America, being the only hyena to do so. Chasmaporthetes managed to survive for some time in North America by deviating from the endurance-running and bone-crushing niches monopolized by canids, and developing into a cheetah-like sprinter. Most of the dog-like hyenas had died off by 1.5 million years ago.

Bone-crushing hyenas
By 10–14 million years ago, the hyena family had split into two distinct groups: dog-like hyenas and bone-crushing hyenas. The arrival of the ancestral bone-crushing hyenas coincided with the decline of the similarly built family Percrocutidae. The bone-crushing hyenas survived the changes in climate and the arrival of canids, which wiped out the dog-like hyenas, though they never crossed into North America, as their niche there had already been taken by the dog subfamily Borophaginae. By 5 million years ago, the bone-crushing hyenas had become the dominant scavengers of Eurasia, primarily feeding on large herbivore carcasses felled by sabre-toothed cats. One genus, Pachycrocuta, was a 200 kg (440 lb) mega-scavenger that could splinter the bones of elephants. Starting in the early Middle Pleistocene Pachycrocuta was replaced by the smaller Crocuta and Hyena, which corresponds to a general faunal change, perhaps in connection to the Mid-Pleistocene transition.

Rise of modern hyenas
The four extant species are the striped hyena (Hyaena hyaena), the brown hyena (Parahyaena brunnea), the spotted hyena (Crocuta crocuta), and the aardwolf (Proteles cristata).

The aardwolf can trace its lineage directly back to Plioviverrops 15 million years ago, and is the only survivor of the dog-like hyena lineage. Its success is partly attributed to its insectivorous diet, for which it faced no competition from canids crossing from North America. It is likely that its unrivaled ability to digest the terpene excretions from soldier termites is a modification of the strong digestive system its ancestors used to consume fetid carrion.

The striped hyena may have evolved from Hyaenictitherium namaquensis of Pliocene Africa. Striped hyena fossils are common in Africa, with records going back as far as the Villafranchian. As fossil striped hyenas are absent from the Mediterranean region, it is likely that the species is a relatively late invader to Eurasia, having likely spread outside Africa only after the extinction of spotted hyenas in Asia at the end of the Ice Age. The striped hyena occurred for some time in Europe during the Pleistocene, having been particularly widespread in France and Germany. It also occurred in Montmaurin, Hollabrunn in Austria, the Furninha Cave in Portugal and the Genista Caves in Gibraltar. The European form was similar in appearance to modern populations, but was larger, being comparable in size to the brown hyena.

The spotted hyena (Crocuta crocuta) diverged from the striped and brown hyena 10 million years ago. Its direct ancestor was the Indian Crocuta sivalensis, which lived during the Villafranchian. Ancestral spotted hyenas probably developed social behaviours in response to increased pressure from rivals on carcasses, thus forcing them to operate in teams. Spotted hyenas evolved sharp carnassials behind their crushing premolars, therefore they did not need to wait for their prey to die, and thus became pack hunters as well as scavengers. They began forming increasingly larger territories, necessitated by the fact that their prey was often migratory, and long chases in a small territory would have caused them to encroach into another clan's turf. Spotted hyenas spread from their original homeland during the Middle Pleistocene, and quickly colonised a very wide area from Europe, to southern Africa and China. The eventual disappearance of the spotted hyena from Europe has traditionally been attributed to the end of the last glacial period and a subsequent displacement of open grassland by closed forests, which favoured wolves and humans instead. However, analyses have shown that climate change alone is insufficient to explain the spotted hyena's disappearance from Europe, suggesting that other factors – such as human pressure – must have played a role. This suggests that the events must be seen within the broader context of late-Quaternary extinctions, as the late Pleistocene and early Holocene saw the disappearance of many primarily large mammals from Europe and the world.

Genera of the Hyaenidae (extinct and recent)
The list follows McKenna and Bell's Classification of Mammals for prehistoric genera (1997) and Wozencraft (2005) in Wilson and Reeders Mammal Species of the World for extant genera. The percrocutids are, in contrast to McKenna and Bell's classification, not included as a subfamily into the Hyaenidae, but as the separate family Percrocutidae (though they are generally grouped as sister-taxa to hyenas ). Furthermore, the living brown hyena and its closest extinct relatives are not included in the genus Pachycrocuta, but in the genus Parahyaena. However, some research has suggested Parahyaena may be synonymous with Pachycrocuta, making the brown hyena the only extant member of this genus.

 * Family Hyaenidae
 * Subfamily Incertae sedis
 * †Tongxinictis (Middle Miocene of Asia)
 * †Subfamily Ictitheriinae
 * †Herpestides (Early Miocene of Africa and Eurasia)
 * †Plioviverrops (including Jordanictis, Protoviverrops, Mesoviverrops; Early Miocene to Early Pliocene of Europe, Late Miocene of Asia)
 * †Ictitherium (=Galeotherium; including Lepthyaena, Sinictitherium, Paraictitherium; Middle Miocene of Africa, Late Miocene to Early Pliocene of Eurasia)
 * †Thalassictis (including Palhyaena, Miohyaena, Hyaenictitherium, Hyaenalopex; Middle to Late Miocene of Asia, Late Miocene of Africa and Europe)
 * †Hyaenotherium (Late Miocene to Early Pliocene of Eurasia)
 * †Miohyaenotherium (Late Miocene of Europe)
 * †Lycyaena (Late Miocene of Eurasia)
 * †Tungurictis (Middle Miocene of Africa and Eurasia)
 * †Protictitherium (Middle Miocene of Africa and Asia, Middle to Late Miocene of Europe)
 * Subfamily Hyaeninae
 * †Palinhyaena (Late Miocene of Asia)
 * †Ikelohyaena (Early Pliocene of Africa)
 * Hyaena (=Euhyaena,=Parahyaena; including striped hyena, Pliohyaena, Pliocrocuta, Anomalopithecus) Early Pliocene (?Middle Miocene) to Recent of Africa, Late Pliocene (?Late Miocene) to Late Pleistocene of Europe, Late Pliocene to recent in Asia
 * Parahyaena (=Hyaena; brown hyena Pliocene to recent of Africa)
 * †Hyaenictis (Late Miocene of Asia?, Late Miocene of Europe, Early Pliocene (?Early Pleistocene) of Africa)
 * †Leecyaena (Late Miocene and/or Early Pliocene of Asia)
 * †Chasmaporthetes (=Ailuriaena; including Lycaenops, Euryboas; Late Miocene to Early Pleistocene of Eurasia, Early Pliocene to Late Pliocene or Early Pleistocene of Africa, Late Pliocene to Early Pleistocene of North America)
 * †Pachycrocuta (Pliocene and Pleistocene of Eurasia and Africa)
 * †Adcrocuta (Late Miocene of Eurasia)
 * Crocuta (=Crocotta; including Eucrocuta; spotted hyena and cave hyena. Late Pliocene to recent of Africa, Late Pliocene to Late Pleistocene of Eurasia)
 * Subfamily Protelinae
 * †Gansuyaena
 * Proteles (=Geocyon; aardwolf. Pleistocene to Recent of Africa)

Phylogeny
The following cladogram illustrates the phylogenetic relationships between extant and extinct hyaenids based on the morphological analysis by Werdelin & Solounias (1991), as updated by Turner et al. (2008).

A more recent molecular analysis agrees on the phylogenetic relationship between the four extant hyaenidae species (Koepfli et al, 2006 ).

Build
Hyenas have relatively short torsos and are fairly massive and wolf-like in build, but have lower hind quarters, high withers and their backs slope noticeably downward towards their rumps. The forelegs are high, while the hind legs are very short and their necks are thick and short. Their skulls superficially resemble those of large canids, but are much larger and heavier, with shorter facial portions. Hyenas are digitigrade, with the fore and hind paws having four digits each and sporting bulging pawpads. Like canids, hyenas have short, blunt, non-retractable claws. Their pelage is sparse and coarse with poorly developed or absent underfur. Most species have a rich mane of long hair running from the withers or from the head. With the exception of the spotted hyena, hyaenids have striped coats, which they likely inherited from their viverrid ancestors. Their ears are large and have simple basal ridges and no marginal bursa. Their vertebral column, including the cervical region are of limited mobility. Hyenas have no baculum. Hyenas have one more pair of ribs than canids do, and their tongues are rough like those of felids and viverrids. Males in most hyena species are larger than females, though the spotted hyena is an exception, as it is the female of the species that outweighs and dominates the male. Also, unlike other hyenas, the female spotted hyena's external genitalia closely resembles that of the male.

Their dentition is similar to that of the canid, but is more specialised for consuming coarse food and crushing bones. The carnassials, especially the upper, are very powerful and are shifted far back to the point of exertion of peak pressure on the jaws. The other teeth, save for the underdeveloped upper molars, are powerful, with broad bases and cutting edges. The canines are short, but thick and robust. Labiolingually, their mandibles are much stronger at the canine teeth than in canids, reflecting the fact that hyenas crack bones with both their anterior dentition and premolars, unlike canids, which do so with their post-carnassial molars. The strength of their jaws is such that both striped and spotted hyenas have been recorded to kill dogs with a single bite to the neck without breaking the skin. The spotted hyena is renowned for its strong bite proportional to its size, but a number of other animals (including the Tasmanian devil) are proportionately stronger. The aardwolf has greatly reduced cheek teeth, sometimes absent in the adult, but otherwise has the same dental formula as the other three species. The dental formula for all hyena species is:

Although hyenas lack perineal scent glands, they have a large pouch of naked skin located at the anal opening. Large anal glands above the anus open into this pouch. Several sebaceous glands are present between the openings of the anal glands and above them. These glands produce a white, creamy secretion that the hyenas paste onto grass stalks. The odor of this secretion is very strong, smelling of boiling cheap soap or burning, and can be detected by humans several meters downwind. The secretions are primarily used for territorial marking, though both the aardwolf and the striped hyena will spray them when attacked.

Behavior
Hyenas groom themselves often like felids and viverrids, and their way of licking their genitals is very cat-like (sitting on the lower back, legs spread with one leg pointing vertically upward). They defecate in the same manner as other Carnivora, though they never raise their legs as canids do when urinating, as urination serves no territorial function for them. Instead, hyenas mark their territories using their anal glands, a trait found also in viverrids and mustelids, but not canids and felids. When attacked by lions or dogs, striped and brown hyenas will feign death, though the spotted hyena will defend itself ferociously. The spotted hyena is very vocal, producing a number of different sounds consisting of whoops, grunts, groans, lows, giggles, yells, growls, laughs and whines. The striped hyena is comparatively silent, its vocalizations being limited to a chattering laugh and howling.

Mating between hyenas involves a number of short copulations with brief intervals, unlike canids, who generally engage in a single, drawn out copulation. Spotted hyena cubs are born almost fully developed, with their eyes open and erupting incisors and canines, though lacking adult markings. In contrast, striped hyena cubs are born with adult markings, closed eyes and small ears. Hyenas do not regurgitate food for their young and male spotted hyenas play no part in raising their cubs, though male striped hyenas do so.

The striped hyena is primarily a scavenger, though it will also attack and kill any animals it can overcome, and will supplement its diet with fruit. The spotted hyena, though it also scavenges occasionally, is an active pack hunter of medium to large sized ungulates, which it catches by wearing them down in long chases and dismembering them in a canid-like manner. Spotted hyenas may kill as many as 95% of the animals they eat. The aardwolf is primarily an insectivore, specialised for feeding on termites of the genus Trinervitermes and Hodotermes, which it consumes by licking them up with its long, broad tongue. An aardwolf can eat 300,000 Trinervitermes on a single outing. Except for the aardwolf, hyenas are known to drive off larger predators, like lions, from their kills, despite having a reputation in popular culture for being cowardly. Hyenas are primarily nocturnal animals, but sometimes venture from their lairs in the early-morning hours. With the exception of the highly social spotted hyena, hyenas are generally not gregarious animals, though the striped and brown hyenas may live in family groups and congregate at kills. Spotted hyenas are one of the few mammals other than bats known to survive infection with rabies virus and have shown little or no disease-induced mortality during outbreaks in sympatric carnivores, in part due to the high concentration of antibodies present in their saliva. Despite this perceived unique disease resistance, little is known about the immune system of spotted hyenas,  and even less is known about other Hyaenidae species.

Folklore, mythology and literature
Spotted hyenas vary in their folkloric and mythological depictions, depending on the ethnic group from which the tales originate. It is often difficult to know whether spotted hyenas are the specific hyena species featured in such stories, particularly in West Africa, as both spotted and striped hyenas are often given the same names. In West African tales, spotted hyenas are sometimes depicted as bad Muslims who challenge the local animism that exists among the Beng in Côte d’Ivoire. In East Africa, Tabwa mythology portrays the spotted hyena as a solar animal that first brought the sun to warm the cold earth, while West African folklore generally shows the hyena as symbolizing immorality, dirty habits, the reversal of normal activities, and other negative traits. In Tanzania, there is a belief that witches use spotted hyenas as mounts. In the Mtwara Region of Tanzania, it is believed that a child born at night while a hyena is crying will likely to grow up to be a thief. In the same area, hyena feces are believed to enable a child to walk at an early age, thus it is not uncommon in that area to see children with hyena dung wrapped in their clothes. The Kaguru of Tanzania and the Kujamaat of southern Senegal view hyenas as inedible and greedy hermaphrodites. A mythical African tribe called the Bouda is reputed to have members able to transform into hyenas. A similar myth occurs in Mansôa. These "werehyenas" are killed when discovered, and do not revert to human form once dead.

Striped hyenas are often referred to in Middle Eastern literature and folklore, typically as symbols of treachery and stupidity. In the Near and Middle East, striped hyenas are generally regarded as physical incarnations of jinns. Arab writer al-Qazwīnī (1204–1283) spoke of a tribe of people called al-Ḍabyūn meaning "hyena people". In his book ‘Ajā’ib Al-Makhlūqāt he wrote that should one of this tribe be in a group of 1,000 people, a hyena could pick him out and eat him. A Persian medical treatise written in 1376 tells how to cure cannibalistic people known as kaftar, who are said to be "half-man, half-hyena". al-Damīrī in his writings in Ḥawayān al-Kubrā (1406) wrote that striped hyenas were vampiric creatures that attacked people at night and sucked the blood from their necks. He also wrote that hyenas only attacked brave people. Arab folklore tells of how hyenas can mesmerise victims with their eyes or sometimes with their pheromones. In a similar vein to al-Damīrī, the Greeks until the end of the 19th century believed that the bodies of werewolves, if not destroyed, would haunt battlefields as vampiric hyenas that drank the blood of dying soldiers. The image of striped hyenas in Afghanistan, India and Palestine is more varied. Though feared, striped hyenas were also symbolic of love and fertility, leading to numerous varieties of love medicine derived from hyena body parts. Among the Baluch and in northern India, witches or magicians are said to ride striped hyenas at night.

The striped hyena is mentioned in the Bible. The Arabic word for the hyena, ḍab` or ḍabu` (plural ḍibā`), is alluded to in a valley in Israel known as Shaqq-ud-Diba` (meaning "cleft of the hyenas") and Wadi-Abu-Diba` (meaning "valley of the hyenas"). Both places have been interpreted by some scholars as being the Biblical Valley of Tsebo`im mentioned in 1 Samuel 13:18. In modern Hebrew, the word for hyena and hypocrite are both the same: tsavua. Though the Authorized King James Version of the Bible interprets the term "`ayit tsavua`" (found in Jeremiah 12:9) as "speckled bird", Henry Baker Tristram argued that it was most likely a hyena being mentioned.

The vocalization of the spotted hyena resembling hysterical human laughter has been alluded to in numerous works of literature: "to laugh like a hyæna" was a common simile, and is featured in The Cobbler's Prophecy (1594), Webster's Duchess of Malfy (1623) and Shakespeare's As You Like It, Act IV. Sc.1.

Die Strandjutwolf (The brown hyena) is an allegorical poem by the renowned South African poet, N. P. van Wyk Louw, which evokes a sinister and ominous presence.

Attacks on humans
In ordinary circumstances, striped hyenas are extremely timid around humans, though they may show bold behaviors towards people at night. On rare occasions, striped hyenas have preyed on humans.

Among hyenas, only the spotted and striped hyenas have been known to become man-eaters. Hyenas are known to have preyed on humans in prehistory: human hair has been found in fossilized hyena dung dating back 195,000 to 257,000 years. Some paleontologists believe that competition and predation by cave hyenas (Crocuta crocuta spelaea) in Siberia was a significant factor in delaying human colonization of Alaska. Hyenas may have occasionally stolen human kills, or entered campsites to drag off the young and weak, much like modern spotted hyenas in Africa. The oldest Alaskan human remains coincide with roughly the same time cave hyenas became extinct, leading some paleontologists to infer that hyena predation prevented humans from crossing the Bering Strait earlier. Hyenas readily scavenge from human corpses; in Ethiopia, hyenas were reported to feed extensively on the corpses of victims of the 1960 attempted coup and the Red Terror. Hyenas habituated to scavenging on human corpses may develop bold behaviors towards living people: hyena attacks on people in southern Sudan increased during the Second Sudanese Civil War, when human corpses were readily available to them.

Although spotted hyenas have been known to prey on humans in modern times, such incidents are rare. However, attacks on humans by spotted hyenas are likely to be underreported. Man-eating spotted hyenas tend to be very large specimens; a pair of man-eating hyenas, responsible for killing 27 people in Mulanje, Malawi in 1962, weighed in at 72 kg (159 lb) and 77 kg (170 lb) after being shot. A 1903 report describes spotted hyenas in the Mzimba district of Angoniland waiting at dawn outside people's huts to attack them when they opened their doors. Victims of spotted hyenas tend to be women, children and sick or infirm men; Theodore Roosevelt wrote in 1908–1909 in Uganda that spotted hyenas regularly killed sufferers of African sleeping sickness as they slept outside in camps. Spotted hyenas are widely feared in Malawi, where they have been known to attack people at night, particularly during the hot season when people sleep outside. A spate of hyena attacks were reported in Malawi's Phalombe plain, with five deaths recorded in 1956, five in 1957 and six in 1958. This pattern continued until 1961, when eight people were killed. Attacks occurred most commonly in September, when people slept outdoors and bush fires made the hunting of wild game difficult for the hyenas. A 2004 news report stated that 35 people were killed by spotted hyenas in a 12-month period in Mozambique along a 20-km stretch of road near the Tanzanian border.

In the 1880s, a hyena was reported to have attacked humans, especially sleeping children, over a three-year period in the Iğdır Province of Turkey, with 25 children and 3 adults being wounded in one year. The attacks provoked local authorities into announcing a reward of 100 rubles for every hyena killed. Further attacks were reported later in some parts of the South Caucasus, particularly in 1908. Instances are known in Azerbaijan of striped hyenas killing children sleeping in courtyards during the 1930s and 1940s. In 1942, a sleeping guard was mauled in his hut by a hyena in Qalıncaq (Golyndzhakh). Cases of children being taken by hyenas by night are known in southeast Turkmenistan's Bathyz Nature Reserve. A further attack on a child was reported around Serakhs in 1948. Several attacks have occurred in India; in 1962, 9 children were thought to have been taken by hyenas in the town of Bhagalpur in the Bihar State in a six-week period, and 19 children up to the age of four were killed by hyenas in Karnataka in 1974. A survey of wild animal attacks during a five-year period in the Indian state of Madhya Pradesh reported that hyenas had attacked three people, causing fewer deaths than wolves, gaur, boar, elephants, tigers, leopards and sloth bears.

Hyenas as food and medicine
Hyenas are used for food and medicinal purposes in Somalia. They are considered halal in Islam. This practice dates back to the times of the Ancient Greeks and Romans, who believed that different parts of the hyena's body were effective means to ward off evil and to ensure love and fertility.

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title: Hyena


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Hershey–Chase experiment

The Hershey–Chase experiments were a series of experiments conducted in 1952 by Alfred Hershey and Martha Chase that helped to confirm that DNA is genetic material. While DNA had been known to biologists since 1869, many scientists still assumed at the time that proteins carried the information for inheritance because DNA appeared to be an inert molecule, and, since it is located in the nucleus, its role was considered to be phosphorus storage. In their experiments, Hershey and Chase showed that when bacteriophages, which are composed of DNA and protein, infect bacteria, their DNA enters the host bacterial cell, but most of their protein does not. Hershey and Chase and subsequent discoveries all served to prove that DNA is the hereditary material.

Hershey shared the 1969 Nobel Prize in Physiology or Medicine with Max Delbrück and Salvador Luria for their "discoveries concerning the genetic structure of viruses".

Historical background
In the early twentieth century, biologists thought that proteins carried genetic information. This was based on the belief that proteins were more complex than DNA. Phoebus Levene's influential "tetranucleotide hypothesis", which incorrectly proposed that DNA was a repeating set of identical nucleotides, supported this conclusion. The results of the Avery–MacLeod–McCarty experiment, published in 1944, suggested that DNA was the genetic material, but there was still some hesitation within the general scientific community to accept this, which set the stage for the Hershey–Chase experiment.

Hershey and Chase, along with others who had done related experiments, confirmed that DNA was the biomolecule that carried genetic information. Before that, Oswald Avery, Colin MacLeod, and Maclyn McCarty had shown that DNA led to the transformation of one strain of Streptococcus pneumoniae to another. The results of these experiments provided evidence that DNA was the biomolecule that carried genetic information.

Methods and results
Hershey and Chase needed to be able to examine different parts of the phages they were studying separately, so they needed to distinguish the phage subsections. Viruses were known to be composed of a protein shell and DNA, so they chose to uniquely label each with a different elemental isotope. This allowed each to be observed and analyzed separately. Since phosphorus is contained in DNA but not amino acids, radioactive phosphorus-32 was used to label the DNA contained in the T2 phage. Radioactive sulfur-35 was used to label the protein sections of the T2 phage, because sulfur is contained in protein but not DNA.

Hershey and Chase inserted the radioactive elements in the bacteriophages by adding the isotopes to separate media within which bacteria were allowed to grow for 4 hours before bacteriophage introduction. When the bacteriophages infected the bacteria, the progeny contained the radioactive isotopes in their structures. This procedure was performed once for the sulfur-labeled phages and once for phosphorus-labeled phages. The labeled progeny were then allowed to infect unlabeled bacteria. The phage coats remained on the outside of the bacteria, while genetic material entered. Disruption of phage from the bacteria by agitation in a blender followed by centrifugation allowed for the separation of the phage coats from the bacteria. These bacteria were lysed to release phage progeny. The progeny of the phages that were labeled with radioactive phosphorus remained labeled, whereas the progeny of the phages labeled with radioactive sulfur were unlabeled. Thus, the Hershey–Chase experiment helped to confirm that DNA, not protein, is the genetic material.

Hershey and Chase showed that the introduction of deoxyribonuclease (referred to as DNase), an enzyme that breaks down DNA, into a solution containing the labeled bacteriophages did not introduce any 32P into the solution. This demonstrated that the phage is resistant to the enzyme while intact. Additionally, they were able to plasmolyze the bacteriophages so that they went into osmotic shock, which effectively created a solution containing most of the 32P and a heavier solution containing structures called "ghosts" that contained the 35S and the protein coat of the virus. It was found that these "ghosts" could adsorb to bacteria that were susceptible to T2, although they contained no DNA and were simply the remains of the original viral capsule. They concluded that the protein protected the DNA from DNase, but that once the two were separated and the phage was inactivated, the DNase could hydrolyze the phage DNA.

Experiment and conclusions
Hershey and Chase were also able to prove that the DNA from the phage is inserted into the bacteria shortly after the virus attaches to its host. Using a high-speed blender they were able to force the bacteriophages from the bacterial cells after adsorption. The lack of 32P-labeled DNA remaining in the solution after the bacteriophages had been allowed to adsorb to the bacteria showed that the phage DNA was transferred into the bacterial cell. The presence of almost all the radioactive 35S in the solution showed that the protein coat that protects the DNA before adsorption stayed outside the cell.

Hershey and Chase concluded that DNA, not protein, was the genetic material. They determined that a protective protein coat was formed around the bacteriophage, but that the internal DNA is what conferred its ability to produce progeny inside a bacterium. They showed that, in growth, protein has no function, while DNA has some function. They determined this from the amount of radioactive material remaining outside of the cell. Only 20% of the 32P remained outside the cell, demonstrating that it was incorporated with DNA in the cell's genetic material. All of the 35S in the protein coats remained outside the cell, showing it was not incorporated into the cell, and that protein is not the genetic material.

Hershey and Chase's experiment concluded that little sulfur-containing material entered the bacterial cell. However no specific conclusions can be made regarding whether material that is sulfur-free enters the bacterial cell after phage adsorption. Further research was necessary to conclude that it was solely bacteriophages' DNA that entered the cell and not a combination of protein and DNA where the protein did not contain any sulfur.

Confirmation
Hershey and Chase concluded that protein was not likely to be the hereditary genetic material. However, they did not make any conclusions regarding the specific function of DNA as hereditary material, and only said that it must have some undefined role.

Confirmation and clarity came a year later in 1953, when James D. Watson and Francis Crick correctly hypothesized, in their journal article "Molecular Structure of Nucleic Acids: A Structure for Deoxyribose Nucleic Acid", the double helix structure of DNA, and suggested the copying mechanism by which DNA functions as hereditary material. Furthermore, Watson and Crick suggested that DNA, the genetic material, is responsible for the synthesis of the thousands of proteins found in cells. They had made this proposal based on the structural similarity that exists between the two macromolecules: both protein and DNA are linear sequences of monomers (amino acids and nucleotides, respectively).

Other experiments
Once the Hershey–Chase experiment was published, the scientific community generally acknowledged that DNA was the genetic code material. This discovery led to a more detailed investigation of DNA to determine its composition as well as its 3D structure. Using X-ray crystallography, the structure of DNA was discovered by James Watson and Francis Crick with the help of previously documented experimental evidence by Maurice Wilkins and Rosalind Franklin. Knowledge of the structure of DNA led scientists to examine the nature of genetic coding and, in turn, understand the process of protein synthesis. George Gamow proposed that the genetic code was composed of sequences of three DNA base pairs known as triplets or codons which represent one of the twenty amino acids. Genetic coding helped researchers to understand the mechanism of gene expression, the process by which information from a gene is used in protein synthesis. Since then, much research has been conducted to modulate steps in the gene expression process. These steps include transcription, RNA splicing, translation, and post-translational modification which are used to control the chemical and structural nature of proteins. Moreover, genetic engineering gives engineers the ability to directly manipulate the genetic materials of organisms using recombinant DNA techniques. The first recombinant DNA molecule was created by Paul Berg in 1972 when he combined DNA from the monkey virus SV40 with that of the lambda phage.

Experiments on hereditary material during the time of the Hershey–Chase experiment often used bacteriophages as a model organism. Bacteriophages lend themselves to experiments on hereditary material because they incorporate their genetic material into their host cell's genetic material (making them useful tools), they multiply quickly, and they are easily collected by researchers.

Legacy
The Hershey–Chase experiment, its predecessors, such as the Avery–MacLeod–McCarty experiment, and successors served to unequivocally establish that hereditary information was carried by DNA. This finding has numerous applications in forensic science, crime investigation and genealogy. It provided the background knowledge for further applications in DNA forensics, where DNA fingerprinting uses data originating from DNA, not protein sources, to deduce genetic variation.

title: Hershey–Chase experiment


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History of religion

The history of religion refers to the written record of human religious feelings, thoughts, and ideas. This period of religious history begins with the invention of writing about 5,200 years ago (3200 BCE). The prehistory of religion involves the study of religious beliefs that existed prior to the advent of written records. One can also study comparative religious chronology through a timeline of religion, or the interrelationships and historical diversification of religious ideologies through the use of evolutionary philosophy and broad comparativism. Writing played a major role in standardizing religious texts regardless of time or location, and making easier the memorization of prayers and divine rules.

The concept of "religion" was formed in the 16th and 17th centuries. Sacred texts like the Bible, the Quran, and others did not have a word or even a concept of religion in the original languages and neither did the people or the cultures in which these sacred texts were written.

The word religion as used in the 21st century does not have an obvious pre-colonial translation into non-European languages. The anthropologist Daniel Dubuisson writes that "what the West and the history of religions in its wake have objectified under the name 'religion' is ... something quite unique, which could be appropriate only to itself and its own history".

History of study
The school of religious history called the Religionsgeschichtliche Schule, a late 19th-century German school of thought, originated the systematic study of religion as a socio-cultural phenomenon. It depicted religion as evolving with human culture, from polytheism to monotheism.

The Religionsgeschichtliche Schule emerged at a time when scholarly study of the Bible and of church history flourished in Germany and elsewhere (see higher criticism, also called the historical-critical method). The study of religion is important: religion and similar concepts have often shaped civilizations' law and moral codes, social structure, art and music.

In order to better understand the origin and current diversity of religious belief systems throughout the world, recent studies have attempted to focus on historical interrelationships and diversification of all major organized religions, applying modern evolutionary philosophy to the comparative analysis of putative ideological groups. These studies take an agnostic, pluralistic approach in the hope of moving beyond chauvinistic cultural tribalism, which is increasingly interfering with our ability to understand "other" cultures and address growing global challenges.

Origin
The earliest archeological evidence interpreted by some as suggestive of the emergence of religious ideas dates back several hundred thousand years, to the Middle and Lower Paleolithic periods: some archaeologists conclude that the apparently intentional burial of archaic humans, Neanderthals and even Homo naledi as early as 300,000 years ago is proof that religious ideas already existed, but such a connection is entirely conjectural. Other evidence that some infer as indicative of religious ideas includes symbolic artifacts from Middle Stone Age sites in Africa. However, the interpretation of early paleolithic artifacts, with regard to how they relate to religious ideas, remains controversial. Archeological evidence from more recent periods is less controversial. Scientists generally interpret a number of artifacts from the Upper Paleolithic (50,000–13,000 BCE) as representing religious ideas. Examples of Upper Paleolithic remains that some associate with religious beliefs include the lion man, the Venus figurines, and the elaborate ritual burial from Sungir.

In the 19th century, researchers proposed various theories regarding the origin of religion, challenging earlier claims of a Christianity-like urreligion. Early theorists, such as Edward Burnett Tylor (1832–1917) and Herbert Spencer (1820–1903), emphasized the concept of animism, while archaeologist John Lubbock (1834–1913) used the term "fetishism". Meanwhile, the religious scholar Max Müller (1823–1900) theorized that religion began in hedonism and the folklorist Wilhelm Mannhardt (1831–1880) suggested that religion began in "naturalism" – by which he meant mythological explanations for natural events. All of these theories have been widely criticized since then; there is no broad consensus regarding the origin of religion.

Pre-pottery Neolithic A (PPNA) Göbekli Tepe, the oldest potentially religious site yet discovered anywhere includes circles of erected massive T-shaped stone pillars, the world's oldest known megaliths decorated with abstract, enigmatic pictograms and carved-animal reliefs. The site, near the home place of original wild wheat, was built before the so-called Neolithic Revolution, i.e., the beginning of agriculture and animal husbandry around 9000 BCE. But the construction of Göbekli Tepe implies organization of an advanced order not hitherto associated with Paleolithic, PPNA, or PPNB societies. The site, abandoned around the time the first agricultural societies started, is still being excavated and analyzed, and thus might shed light on the significance it had, if any, for the religions of older, foraging communities, as well as for the general history of religions.

The Pyramid Texts from ancient Egypt, the oldest known religious texts in the world, date to between 2400 and 2300 BCE.

The earliest records of Indian religion are the Vedas, composed c. 1500–1200 BCE during the Vedic Period.

Surviving early copies of religious texts include:

 * The Upanishads, some of which date to the mid-first millennium BCE.
 * The Dead Sea Scrolls, representing fragmentary texts of the Hebrew Tanakh.
 * Complete Hebrew texts, also of the Tanakh, but translated into the Greek language (Septuagint 300–200 BCE), were in wide use by the early 1st century CE.
 * The Zoroastrian Avesta, from a Sassanian-era master copy.

Axial age
Some historians have labelled the period from 900 to 200 BCE as the "axial age", a term coined by German-Swiss philosopher Karl Jaspers (1883–1969). According to Jaspers, in this era of history "the spiritual foundations of humanity were laid simultaneously and independently... And these are the foundations upon which humanity still subsists today." Intellectual historian Peter Watson has summarized this period as the foundation time of many of humanity's most influential philosophical traditions, including monotheism in Persia and Canaan, Platonism in Greece, Buddhism and Jainism in India, and Confucianism and Taoism in China. These ideas would become institutionalized in time – note for example Ashoka's role in the spread of Buddhism, or the role of Neoplatonic philosophy in Christianity at its foundation.

The historical roots of Jainism in India date back to the 9th century BCE with the rise of Parshvanatha and his non-violent philosophy.

Middle Ages
World religions of the present day established themselves throughout Eurasia during the Middle Ages by:

 * Christianization of the Western world
 * Buddhist missions to East Asia
 * the decline of Buddhism in the Indian subcontinent
 * the spread of Islam throughout the Middle East, Central Asia, North Africa and parts of Europe and India

During the Middle Ages, Muslims came into conflict with Zoroastrians during the Muslim conquest of Persia (633–654); Christians fought against Muslims during the Arab–Byzantine wars (7th to 11th centuries), the Crusades (1095 onward), the Reconquista (718–1492), the Ottoman wars in Europe (13th century onwards) and the Inquisition; Shamanism was in conflict with Buddhists, Taoists, Muslims and Christians during the Mongol invasions and conquests (1206–1337); and Muslims clashed with Hindus and Sikhs during the Muslim conquests in the Indian subcontinent (8th to 16th centuries).

Many medieval religious movements continued to emphasize mysticism, such as the Cathars and related movements in the West, the Jews in Spain (see Zohar), the Bhakti movement in India and Sufism in Islam. Monotheism and related mysticisms reached definite forms in Christian Christology and in Islamic Tawhid. Hindu monotheist notions of Brahman likewise reached their classical form with the teaching of Adi Shankara (788–820).

Modern Ages
From the 15th century to the 19th century, European colonisation resulted in the spread of Christianity to Sub-Saharan Africa, and the Americas, Australia and the Philippines. The invention of the printing press in the 15th century played a major role in the rapid spread of the Protestant Reformation under leaders such as Martin Luther (1483–1546) and John Calvin (1509–1564). Wars of religion broke out, culminating in the Thirty Years' War which ravaged Central Europe between 1618 and 1648. The 18th century saw the beginning of secularisation in Europe, a trend which gained momentum after the French Revolution broke out in 1789. By the late 20th century, religion had declined in most of Europe.

Shamanism and ancestor worship

 * Prehistoric religion
 * Shamanism
 * Animism
 * Ancestor worship
 * Tribal religion

Panentheism

 * Sikhism
 * Neoplatonism
 * Eastern Orthodoxy

Polytheism

 * Ancient Near Eastern religion, Egyptian mythology
 * Ancient Greek religion, Ancient Roman religion
 * Germanic paganism, Finnish Paganism, Norse paganism
 * Maya religion, Inca religion, Aztec religion
 * Neopaganism, Polytheistic reconstructionism

Monotheism

 * Aten
 * Baháʼí Faith
 * History of the Baháʼí Faith
 * Judaism
 * History of Mandaeism
 * Neoplatonism
 * History of Christianity
 * History of the Catholic Church
 * History of Protestantism
 * History of Jehovah's Witnesses
 * Mormonism
 * History of the Latter Day Saint movement
 * History of The Church of Jesus Christ of Latter-day Saints
 * History of Islam
 * Zoroastrianism

Monism

 * History of Buddhism
 * History of Jainism
 * History of Hinduism

Dualism

 * Gnosticism

New religious movements

 * Rastafari movement
 * History of Wicca
 * Timeline of Scientology
 * Adventism
 * Jehovah's Witnesses
 * Mormonism
 * Pentecostalism
 * Bábism
 * Bahá'í Faith
 * Spiritualism
 * History of Spiritism
 * Thelema
 * Ahmadiyya
 * Nation of Islam

title: History of religion