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Economy of Cyprus

The economy of Cyprus is a high-income economy as classified by the World Bank, and was included by the International Monetary Fund in its list of advanced economies in 2001. Cyprus adopted the euro as its official currency on 1 January 2008, replacing the Cypriot pound at an irrevocable fixed exchange rate of CYP 0.585274 per €1.

The 2012–2013 Cypriot financial crisis, part of the wider European debt crisis, has dominated the country's economic affairs in recent times. In March 2013, the Cypriot government reached an agreement with its eurozone partners to split the country's second biggest bank, the Cyprus Popular Bank (also known as Laiki Bank), into a "bad" bank which would be wound down over time and a "good" bank which would be absorbed by the larger Bank of Cyprus. In return for a €10 billion bailout from the European Commission, the European Central Bank and the International Monetary Fund, the Cypriot government would be required to impose a significant haircut on uninsured deposits. Insured deposits of €100,000 or less would not be affected. After a three-and-a-half-year recession, Cyprus returned to growth in the first quarter of 2015. Cyprus successfully concluded its three-year financial assistance programme at the end of March 2016, having borrowed a total of €6.3 billion from the European Stability Mechanism and €1 billion from the IMF. The remaining €2.7 billion of the ESM bailout was never dispensed, due to the Cypriot government's better than expected finances over the course of the programme.

Economy in the government-controlled area
Cyprus has an open, free-market, service-based economy with some light manufacturing. Internationally, Cyprus promotes its geographical location as a "bridge" between East and West, along with its educated English-speaking population, moderate local costs, good airline connections, and telecommunications.

Since gaining independence from the United Kingdom in 1960, Cyprus has had a record of successful economic performance, reflected in strong growth, full employment conditions and relative stability. The underdeveloped agrarian economy inherited from colonial rule has been transformed into a modern economy, with dynamic services, industrial and agricultural sectors and an advanced physical and social infrastructure. The Cypriots are among the most prosperous people in the Mediterranean region, with GDP per capita in 2023 exceeding $37,000 in nominal terms and approaching $59,000 on the basis of purchasing power parity.

Their standard of living is reflected in the country's "very high" Human Development Index, by which it ranks 29th out of 191 countries in the world.

However, after more than three decades of unbroken growth, the Cypriot economy contracted in 2009. This reflected the exposure of Cyprus to the Great Recession and European debt crisis. Furthermore, Cyprus was dealt a severe blow by the Evangelos Florakis Naval Base explosion in July 2011, with the cost to the economy estimated at €1–3 billion, or up to 17% of GDP.

The economic achievements of Cyprus during the preceding decades have been significant, bearing in mind the severe economic and social dislocation created by the Turkish invasion of 1974 and the continuing occupation of the northern part of the island by Turkey. The Turkish invasion inflicted a serious blow to the Cyprus economy and in particular to agriculture, tourism, mining and Quarrying: 70 percent of the island's wealth-producing resources were lost, the tourist industry lost 65 percent of its hotels and tourist accommodation, the industrial sector lost 46 percent, and mining and quarrying lost 56 percent of production. The loss of the port of Famagusta, which handled 83 percent of the general cargo, and the closure of Nicosia International Airport, in the buffer zone, were additional setbacks.

The success of Cyprus in the economic sphere has been attributed, inter alia, to the adoption of a market-oriented economic system, the pursuance of sound macroeconomic policies by the government as well as the existence of a dynamic and flexible entrepreneurship and a highly educated labor force. Moreover, the economy benefited from the close cooperation between the public and private sectors.

In the past 30 years, the economy has shifted from agriculture to light manufacturing and services. The services sector, including tourism, contributes almost 80% to GDP and employs more than 70% of the labor force. Industry and construction account for approximately one-fifth of GDP and labor, while agriculture is responsible for 2.1% of GDP and 8.5% of the labor force. Potatoes and citrus are the principal export crops. After robust growth rates in the 1980s (average annual growth was 6.1%), economic performance in the 1990s was mixed: real GDP growth was 9.7% in 1992, 1.7% in 1993, 6.0% in 1994, 6.0% in 1995, 1.9% in 1996 and 2.3% in 1997. This pattern underlined the economy's vulnerability to swings in tourist arrivals (i.e., to economic and political conditions in Cyprus, Western Europe, and the Middle East) and the need to diversify the economy. Declining competitiveness in tourism and especially in manufacturing are expected to act as a drag on growth until structural changes are effected. Overvaluation of the Cypriot pound prior to the adoption of the euro in 2008 had kept inflation in check.

Trade is vital to the Cypriot economy — the island is not self-sufficient in food and until the recent offshore gas discoveries had few known natural resources – and the trade deficit continues to grow. Cyprus must import fuels, most raw materials, heavy machinery, and transportation equipment. More than 50% of its trade is with the rest of the European Union, especially Greece and the United Kingdom, while the Middle East receives 20% of exports. In 1991, Cyprus introduced a value-added tax (VAT), which is at 19% as of 13 January 2014. Cyprus ratified the new world trade agreement (General Agreement on Tariffs and Trade, GATT) in 1995 and began implementing it fully on 1 January 1996. EU accession negotiations started on 31 March 1998, and concluded when Cyprus joined the organization as a full member in 2004.

Investment climate
The Cyprus legal system is founded on English law, and is therefore familiar to most international financiers. Cyprus's legislation was aligned with EU norms in the period leading up to EU accession in 2004. Restrictions on foreign direct investment were removed, permitting 100% foreign ownership in many cases. Foreign portfolio investment in the Cyprus Stock Exchange was also liberalized. In 2002 a modern, business-friendly tax system was put in place with a 12.5% corporate tax rate, one of the lowest in the EU. Cyprus has concluded treaties on double taxation with more than 40 countries, and, as a member of the Eurozone, has no exchange restrictions. Non-residents and foreign investors may freely repatriate proceeds from investments in Cyprus.

Role as a financial hub
In the years following the dissolution of the Soviet Union it gained great popularity as a portal for investment from the West into Russia and Eastern Europe, becoming for companies of that origin the most common tax haven. More recently, there have been increasing investment flows from the West through Cyprus into Asia, particularly China and India, South America and the Middle East. In addition, businesses from outside the EU use Cyprus as their entry-point for investment into Europe. The business services sector remains the fastest growing sector of the economy, and had overtaken all other sectors in importance. CIPA has been fundamental towards this trend.

Following the 2022 Russian invasion of Ukraine, Cyprus businesses and individuals have come under scrutiny and criticism for allowing EU and US sanctions to be breached with belated attempts to stop them or bring the culprits to justice. A number of professional law and accounting firms have been identified as helping Russian Oligarchs evade sanctions.

In January 2024, during a Council of Europe Parliamentary Assembly, Cypriot President Nikos Christodoulides maintained that Cyrprus had fully implemented all European Union sanctions against Russia. Around the same time of his speech, it was reported that Russian deposits held in Cypriot banks had fallen 76% from 2014 to 2022. The number of Russian clients using Cypriot banks also dropped 82% in the same period.

Agriculture
Cyprus produced in 2018:

 * 106 thousand tons of potato;
 * 37 thousand tons of tangerine;
 * 23 thousand tons of grape;
 * 20 thousand tons of orange;
 * 19 thousand tons of grapefruit;
 * 19 thousand tons of olive;
 * 18 thousand tons of wheat;
 * 18 thousand tons of barley;
 * 15 thousand tons of tomato;
 * 13 thousand tons of watermelon;
 * 10 thousand tons of melon;

In addition to smaller productions of other agricultural products.

Oil and gas
Surveys suggest more than 100 trillion cubic feet (2.831 trillion cubic metres) of reserves lie untapped in the eastern Mediterranean basin between Cyprus and Israel – almost equal to the world's total annual consumption of natural gas. In 2011, Noble Energy estimated that a pipeline to Leviathan gas field could be in operation as soon as 2014 or 2015. In January 2012, Noble Energy announced a natural gas field discovery. It attracted Shell, Delek and Avner as partners. Several production sharing contracts for exploration were signed with international companies, including Eni, KOGAS, TotalEnergies, ExxonMobil and QatarEnergy. It is necessary to develop infrastructure for landing the gas in Cyprus and for liquefaction for export.

Role as a shipping hub
Cyprus constitutes one of the largest ship management centers in the world; around 50 ship management companies and marine-related foreign enterprises are conducting their international activities in the country while the majority of the largest ship management companies in the world have established fully fledged offices on the island. Its geographical position at the crossroads of three continents and its proximity to the Suez Canal has promoted merchant shipping as an important industry for the island nation. Cyprus has the tenth-largest registered fleet in the world, with 1,030 vessels accounting for 31,706,000 dwt as of 1 January 2013.

Tourism
Tourism is an important factor of the island state's economy, culture, and overall brand development. With over 2 million tourist arrivals per year, it is the 40th most popular destination in the world. However, per capita of local population, it ranks 17th. The industry has been honored with various international awards, spanning from the Sustainable Destinations Global Top 100, VISION on Sustainable Tourism, Totem Tourism and Green Destination titles bestowed to Limassol and Paphos in December 2014. The island beaches have been awarded with 57 Blue Flags. Cyprus became a full member of the World Tourism Organization when it was created in 1975. According to the World Economic Forum's 2013 Travel and Tourism Competitiveness Index, Cyprus' tourism industry ranks 29th in the world in terms of overall competitiveness. In terms of Tourism Infrastructure, in relation to the tourism industry Cyprus ranks 1st in the world. The Cyprus Tourism Organization has a status of a semi-governmental organisation charged with overseeing the industry practices and promoting the island worldwide.

Trade
In 2008 fiscal aggregate value of goods and services exported by Cyprus was in the region of $1.53 billion. It primarily exported goods and services such as citrus fruits, cement, potatoes, clothing and pharmaceuticals. At that same period total financial value of goods and services imported by Cyprus was about $8.689 billion. Prominent goods and services imported by Cyprus in 2008 were consumer goods, machinery, petroleum and other lubricants, transport equipment and intermediate goods.

Cypriot trade partners
Traditionally Greece has been a major export and import partner of Cyprus. In fiscal 2007, it amounted for 21.1 percent of total exports of Cyprus. At that same period it was responsible for 17.7 percent of goods and services imported by Cyprus. Some other important names in this regard are UK and Italy.

Eurozone crisis
In 2012, Cyprus became affected by the Eurozone financial and banking crisis. In June 2012, the Cypriot government announced it would need €1.8 billion of foreign aid to support the Cyprus Popular Bank, and this was followed by Fitch down-grading Cyprus's credit rating to junk status. Fitch said Cyprus would need an additional €4 billion to support its banks and the downgrade was mainly due to the exposure of Bank of Cyprus, Cyprus Popular Bank and Hellenic Bank (Cyprus's 3 largest banks) to the Greek financial crisis.

In June 2012 the Cypriot finance minister, Vassos Shiarly, stated that the European Central Bank, European commission and IMF officials are to carry out an in-depth investigation into Cyprus' economy and banking sector to assess the level of funding it requires. The Ministry of Finance rejected the possibility that Cyprus would be forced to undergo the sweeping austerity measures that have caused turbulence in Greece, but admitted that there would be "some negative repercussion".

In November 2012 international lenders negotiating a bailout with the Cypriot government have agreed on a key capital ratio for banks and a system for the sector's supervision. Both commercial banks and cooperatives will be overseen by the Central Bank and the Ministry of Finance. They also set a core Tier 1 ratio – a measure of financial strength – of 9% by the end of 2013 for banks, which could then rise to 10% in 2014.

In 2014, Harris Georgiades pointed that exiting the Memorandum with the European troika required a return to the markets. This he said, required "timely, effective and full implementation of the program." The Finance Minister stressed the need to implement the Memorandum of understanding without an additional loan.

In 2015, Cyprus was praised by the President of the European Commission for adopting the austerity measures and not hesitating to follow a tough reform program.

In 2016, Moody's Investors Service changed its outlook on the Cypriot banking system to positive from stable, reflecting the view that the recovery will restore banks to profitability and improve asset quality. The quick economic recovery was driven by tourism, business services and increased consumer spending. Creditor confidence was also strengthened, allowing Bank of Cyprus to reduce its Emergency Liquidity Assistance to €2.0 billion (from €9.4 billion in 2013). Within the same period, Bank of Cyprus chairman Josef Ackermann urged the European Union to pledge financial support for a permanent solution to the Cyprus dispute.

Economy of Northern Cyprus
The economy of Turkish-occupied northern Cyprus is about one-fifth the size of the economy of the government-controlled area, while GDP per capita is around half. Because the de facto administration is recognized only by Turkey, it has had much difficulty arranging foreign financing, and foreign firms have hesitated to invest there. The economy mainly revolves around the agricultural sector and government service, which together employ about half of the work force.

The tourism sector also contributes substantially into the economy. Moreover, the small economy has seen some downfalls because the Turkish lira is legal tender. To compensate for the economy's weakness, Turkey has been known to provide significant financial aid. In both parts of the island, water shortage is a growing problem, and several desalination plants are planned.

The economic disparity between the two communities is pronounced. Although the economy operates on a free-market basis, the lack of private and government investment, shortages of skilled labor and experienced managers, and inflation and the devaluation of the Turkish lira continue to plague the economy.

Trade with Turkey
Turkey is by far the main trading partner of Northern Cyprus, supplying 55% of imports and absorbing 48% of exports. In a landmark case, the European Court of Justice (ECJ) ruled on 5 July 1994 against the British practice of importing produce from Northern Cyprus based on certificates of origin and phytosanitary certificates granted by the de facto authorities. The ECJ decided that only goods bearing certificates of origin from the internationally recognized Republic of Cyprus could be imported by EU member states. The decision resulted in a considerable decrease of Turkish Cypriot exports to the EU: from $36.4 million (or 66.7% of total Turkish Cypriot exports) in 1993 to $24.7 million in 1996 (or 35% of total exports) in 1996. Even so, the EU continues to be the second-largest trading partner of Northern Cyprus, with a 24.7% share of total imports and 35% share of total exports.

The most important exports of Northern Cyprus are citrus and dairy products. These are followed by rakı, scrap and clothing.

Assistance from Turkey is the mainstay of the Turkish Cypriot economy. Under the latest economic protocol (signed 3 January 1997), Turkey has undertaken to provide loans totalling $250 million for the purpose of implementing projects included in the protocol related to public finance, tourism, banking, and privatization. Fluctuation in the Turkish lira, which suffered from hyperinflation every year until its replacement by the Turkish new lira in 2005, exerted downward pressure on the Turkish Cypriot standard of living for many years.

The de facto authorities have instituted a free market in foreign exchange and permit residents to hold foreign-currency denominated bank accounts. This encourages transfers from Turkish Cypriots living abroad.

Happiness
Economic factors such as the GDP and national income strongly correlate with the happiness of a nation's citizens. In a study published in 2005, citizens from a sample of countries were asked to rate how happy or unhappy they were as a whole on a scale of 1 to 7 (Ranking: 1. Completely happy, 2. Very happy, 3. Fairly happy,4. Neither happy nor unhappy, 5. Fairly unhappy, 6. Very unhappy, 7. Completely unhappy.) Cyprus had a score of 5.29. On the question of how satisfied citizens were with their main job, Cyprus scored 5.36 on a scale of 1 to 7 (Ranking: 1. Completely satisfied, 2. Very satisfied, 3. Fairly satisfied, 4. Neither satisfied nor dissatisfied, 5. Fairly dissatisfied, 6. Very dissatisfied, 7. Completely dissatisfied.) In another ranking of happiness, Northern Cyprus ranks 58 and Cyprus ranks 61, according to the 2018 World Happiness Report. The report rates 156 countries based on variables including income, healthy life expectancy, social support, freedom, trust, and generosity.

Economic factors play a significant role in the general life satisfaction of Cyprus citizens, especially with women who participate in the labor force at a lower rate, work in lower ranks, and work in more public and service sector jobs than the men. Women of different skill-sets and "differing economic objectives and constraints" participate in the tourism industry. Women participate in this industry through jobs like hotel work to serve and/or bring pride to their family, not necessarily to satisfy their own selves. In this study, women with income higher than the mean household income reported higher levels of satisfaction with their lives while those with lower income reported the opposite. When asked who they compare themselves with (those with lower, same, or higher economic status), results showed that those that compared themselves with people of higher economic statuses than them had the lowest level of life satisfaction. While the correlation of income and happiness is positive, it is significantly low; there is stronger correlation between comparison and happiness. This indicates that not only income level but income level in relation to that of others affects their amount of life satisfaction.

Classified as a Mediterranean welfare regime, Cyprus has a weak public Welfare system. This means there is a strong reliance on the family, instead of the state, for both familial and economic support. Another finding is that being a full-time housewife has a stronger negative effect on happiness for women of Northern Cyprus than being unemployed, showing how the combination of gender and the economic factor of participating in the labor force affects life satisfaction. Economic factors also negatively correlate with the happiness levels of those that live in the capital city: citizens living in the capital express lower levels of happiness. As found in this study, citizens of Cyprus that live in its capital, Nicosia, are significantly less happy than others whether or not socio-economic variables are controlled for. Another finding was that the young people in the capital are unhappier than the rest of Cyprus; the old are not.

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Transport in Cyprus

Transport in Cyprus consists of transport by land, water and air. Road transport is the primary mode of transport for most Cypriot citizens, and Cyprus's road transport systems are well-developed and extensively used across the island.

Because Cyprus no longer has a working railway system, various other methods of transport are needed to ensure the proper delivery of any cargo, be it human or freight. As the last passenger railway was dismantled in 1952, the only remaining modes of transport are by road, sea, and air.

Roads

From the 12,118 km of roads in the areas controlled by the Republic of Cyprus in 2006, 7,850 km were paved, while 4,268 km were unpaved. In 1996, the Turkish Cypriot area showed a close, but smaller ratio of paved to unpaved with about 1,370 km out of 2,350 km paved and 980 km unpaved. As a legacy of British rule, Cyprus is one of only three EU nations in which vehicles drive on the left.

Motorways

 * A1 Nicosia to Limassol
 * A2 connects A1 near Pera Chorio with A3 by Larnaca
 * A3 Larnaca Airport to Agia Napa, also serves as a circular road for Larnaca.
 * A5 connects A1 near Kofinou with A3 by Larnaca
 * A6 Pafos to Limassol
 * A7 Pafos to Polis (final plans)
 * A9 Nicosia to Astromeritis
 * A22 Dali industrial area to Anthoupolis, Lakatamia (Nicosia 3rd ring road, final plans)

Public transportation
Nicosia's residents rely on private cars to go around the city. With more than 629 automobiles per 1,000 people, Cyprus has one of the highest car ownership rates in the world and the country uses very little public transportation. Only 3% of journeys in the Greater Nicosia urban region are made by public transportation. Cycling is considerably less common at 2%. The government of Cyprus and authorities of Nicosia have developed a public transportation plan to ensure access to more areas and provide more options, apart from private cars.

Public transportation companies
In Cyprus, public transportation by bus is run by different companies based on the district.

Nicosia and Larnaca: NPT (Nicosia Public Transport) and LPT (Larnaca Public Transport), operated by Cyprus Public Transport (CPT)

 Limassol:  EMEL (Transport Company for Limassol Commuters)

 Paphos:  OSYPA (Paphos Transport Organisation)

 Famagusta:  OSEA (Famagusta District Transport Organisation)

 INTERCITY BUSES:  Services transport between all major cities

Public buses
In 2006, extensive plans were announced to improve and expand bus services and restructure public transport throughout Cyprus, with the financial backing of the European Union Development Bank. In 2010, the new revised and expanded bus network was implemented into the system.

In 2020, the transport companies for the districts of Nicosia and Larnaca were changed from OSEL (Nicosia District Transport Organisation) to NPT (Nicosia Public Transport) and from ZENON Larnaca Buses to LPT (Larnaca Public Transport) respectively.

In 2022, Cyprus Public Transport made new plans for Nicosia's Public Transport by changing route numbers, adding new bus hubs and modernising buses and the all-out feel of the transport system. The plan has been introduced in two phases and is currently completed.

Rail
Cyprus currently has no functioning railway systems. The last of the narrow gauge systems in the country closed in 1974. There had been studies and preparatory work done to establish a modern system between the major cities, motivated by worsening traffic issues. But as of 2024, there has been no funding commitment towards a project.

In 2018, Nicosia municipal authorities requested an opinion on the construction of a tram network in the city to the European Investment Bank's JASPERS strategists, who concluded a need for a phased approach. Relevant studies were also carried out in the past, but no concrete plans has materialised.

Licensed vehicles
Road transport is the dominant form of transport on the island. Figures released by the International Road Federation in 2007 show that Cyprus holds the highest car ownership rate in the world with 742 cars per 1,000 people.

Public transport in Cyprus is limited to privately run bus services (except in Nicosia and Larnaca), taxis, and interurban 'shared' taxi services (locally referred to as service taxis). Thus, private car ownership in the country is the fifth highest per capita in the world. However, in 2006 extensive plans were announced to expand and improve bus services and restructure public transport throughout Cyprus, with the financial backing of the European Union Development Bank

Sea harbours and ports
The ports of Cyprus are operated and maintained by the Cyprus Ports Authority. Major harbours of the island are Limassol Harbour, and Larnaca Harbour, which service cargo, passenger, and cruise ships. Limassol is the larger of the two, and handles a large volume of both cargo and cruise vessels. Larnaca is primarily a cargo port but played a big part in the evacuation of foreign nationals from Lebanon in 2006, and in the subsequent humanitarian aid effort. A smaller cargo dock also exists at Vasilikos, near Zygi (a small town between Larnaca and Limassol). Smaller vessels and private yachts can dock at Marinas in Cyprus.

Public bicycle sharing system

Nextbike is the latest transportation system in Cyprus, similar to programs employed successfully in various cities around the world. Bicycles can be found at stations in Nicosia and Limassol, as well as with 1 station in Larnaca.

Merchant marine

 * See full article on Cyprus Merchant Marine

Total: 1,414 ships (with a volume of or over) totaling /

Ships by Type: barge carrier 2, bulk carrier 442, cargo ship 495, chemical tanker 22, combination bulk 40, combination ore/oil 8, container ship 144, Liquified Gas Carrier 6, passenger ship 8, petroleum tanker 142, refrigerated cargo 41, roll-on/roll-off 45, short-sea passenger 13, specialized tanker 4, vehicle carrier 2 (1999 est.)

Airports
In 1999, Cyprus had 12 airports with paved runways. Of them, seven had runways of lengths between 2,438 and 3,047 metres, one had a length between 1,524 and 2,437 metres, three had lengths between 914 and 1524 metres, and one had a length less than 914 metres.

Of the three airports with unpaved runways, two had lengths less than 914 metres and one had a length between 914 and 1524 metres.

International airports
Larnaca International Airport is the island's main airport and flies to many locations worldwide.

Paphos International Airport is the 2nd largest airport and mostly flies to Europe, via Ryanair; with occasional flights to other continents.

Nicosia International Airport is an abandoned airport. It used to be the island's main airport until 1974. It remains closed to the public.

Ercan International Airport is the main airport in the de facto state of Northern Cyprus. The airport's only destination is Turkey, serviced only by Turkish airlines (not to be confused with the company). Flights to and from Ercan Airport are illegal.

license: Creative Commons - Attribution Share-Alike - https://creativecommons.org/licenses/by-sa/4.0/

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Foreign relations of Cyprus

Cyprus is a member of the United Nations along with most of its agencies as well as the Commonwealth of Nations, World Bank, International Monetary Fund and Council of Europe. In addition, the country has signed the General Agreement on Tariffs and Trade (GATT) and the Multilateral Investment Guarantee Agency Agreement (MIGA). Cyprus has been a member of the European Union since 2004 and in the second half of 2012 it held the Presidency of the Council of the European Union.

Historical non-alignment
Cyprus has historically followed a non-aligned foreign policy, although it increasingly identifies with the West in its cultural affinities and trade patterns, and maintains close relations with the European Union, Greece, Armenia, Israel and the United States.

The prime originator of Cypriot non-alignment was Archbishop of Cyprus Makarios III, the first President (1960–1977) of the independent republic of Cyprus. Prior to independence, Makarios - by virtue of his post as Archbishop of Cyprus and head of the Cypriot Orthodox Church - was the Greek Cypriot Ethnarch, or de facto leader of the community. A highly influential figure well before independence, he participated in the 1955 Bandung Conference. After independence, Makarios took part in the 1961 founding meeting of the Non-Aligned Movement in Belgrade.

Reasons for this neutrality may lie in the extreme pressures exerted on the infant Republic by its larger neighbours, Turkey and Greece. Intercommunal rivalries and movements for union with Greece or partial union with Turkey may have persuaded Makarios to steer clear of close affiliation with either side. In any case Cyprus became a high-profile member of the Non-Aligned Movement and retained its membership until its entry into the European Union in 2004. At the non-governmental level, Cyprus has also been a member of the popular extension of the Non-Aligned Movement, the Afro-Asian Peoples' Solidarity Organisation hosting several high-level meetings.

Immediately after the 1974 Greek-sponsored coup d'état and the Turkish invasion, Makarios secured international recognition of his administration as the legitimate government of the whole island. This was disputed only by Turkey, which currently recognizes only the Turkish Republic of Northern Cyprus, established in 1983.

Since the 1974 crisis, the chief aim of the foreign policy of the Republic of Cyprus has been to secure the withdrawal of Turkish forces and the reunification of the island under the most favorable constitutional and territorial settlement possible. This campaign has been pursued primarily through international forums such as the United Nations and the Non-Aligned Movement, and in recent years through the European Union.

Diplomatic relations
List of countries which Cyprus maintains diplomatic relations with:

Europe
Cyprus' 1990 application for full EU membership caused a storm in the Turkish Cypriot community, which argued that the move required their consent. Following the December 1997 EU Summit decisions on EU enlargement, accession negotiations began 31 March 1998. Cyprus joined the European Union on 1 May 2004. To fulfil its commitment as a member of the European Union, Cyprus withdrew from the Non-Aligned Movement on accession, retaining observer status.

Multilateral relations

 * Cyprus–NATO relations

Overview
The Republic of Cyprus maintains diplomatic relations with 179 states (including the Holy See and Palestinian National Authority) and is United Nations, Union for the Mediterranean and European Union full member. It does not maintain diplomatic relations with:
 * 🇦🇿 Azerbaijan
 * Republic of Congo, 🇨🇫 Central African Republic, 🇩🇯 Djibouti
 * 🇧🇹 Bhutan
 * 🇰🇮 Kiribati
 * 🇭🇹 Haiti

The Republic of Cyprus is not recognised by 🇹🇷 Turkey.

International disputes
The 1974 invasion of the Turkish army divided the island nation into two. The internationally recognised Republic of Cyprus currently has effective control in the south of the island (59% of the island's land area) while its area not under its effective control makes up 37% of the island. Turkey utilising the territory occupied during the invasion recognizes a declared separatist UDI of Turkish Cypriots in 1983, contrary to multiple United Nations Security Council Resolutions. The two territories of the Republic are separated by a United Nations Buffer Zone (4% of the island); there are two UK sovereign base areas mostly within the Greek Cypriot portion of the island.

title: Foreign relations of Cyprus


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Talk:Cyprus

Semi-protected edit request on 9 May 2024
Grammar correction: In the History-Prehistoric and Ancient Cyprus section towards the end, "Cypriot fleet were also sent to help Amphoterus." should be "The Cypriot fleet was also sent to help Amphoterus." Cherryfresca (talk) 18:34, 9 May 2024 (UTC)
 * ✅ – dudhhr talkcontribssheher 18:37, 9 May 2024 (UTC)

Semi-protected edit request on 19 June 2024
Remove or relocate the first sentence about water wells? Seems very out of place 208.97.202.135 (talk) 22:12, 19 June 2024 (UTC)
 * Red question icon with gradient background.svg Not done: it's not clear what changes you want to be made. Please mention the specific changes in a "change X to Y" format and provide a reliable source if appropriate. M.Bitton (talk) 16:56, 20 June 2024 (UTC)

Why is there no mention of the Latins in Cyprus?
Why is there not mention of the Latins as an Ethnic Minority Group section? Maximos2017 (talk) 10:44, 2 July 2024 (UTC)

title: Talk:Cyprus


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Cretaceous

The Cretaceous is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin creta, "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation Kreide.

The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was largely ice-free, although there is some evidence of brief periods of glaciation during the cooler first half, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth by the end of the Cretaceous, coincident with the decline and extinction of previously widespread gymnosperm groups.

The Cretaceous (along with the Mesozoic) ended with the Cretaceous–Paleogene extinction event, a large mass extinction in which many groups, including non-avian dinosaurs, pterosaurs, and large marine reptiles, died out, widely thought to have been caused by the impact of a large asteroid that formed the Chicxulub crater in the Gulf of Mexico. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary (K–Pg boundary), a geologic signature associated with the mass extinction that lies between the Mesozoic and Cenozoic Eras.

Etymology and history
The Cretaceous as a separate period was first defined by Belgian geologist Jean d'Omalius d'Halloy in 1822 as the Terrain Crétacé, using strata in the Paris Basin and named for the extensive beds of chalk (calcium carbonate deposited by the shells of marine invertebrates, principally coccoliths), found in the upper Cretaceous of Western Europe. The name Cretaceous was derived from the Latin creta, meaning chalk. The twofold division of the Cretaceous was implemented by Conybeare and Phillips in 1822. Alcide d'Orbigny in 1840 divided the French Cretaceous into five étages (stages): the Neocomian, Aptian, Albian, Turonian, and Senonian, later adding the Urgonian between Neocomian and Aptian and the Cenomanian between the Albian and Turonian.

Subdivisions
The Cretaceous is divided into Early and Late Cretaceous epochs, or Lower and Upper Cretaceous series. In older literature, the Cretaceous is sometimes divided into three series: Neocomian (lower/early), Gallic (middle) and Senonian (upper/late). A subdivision into 12 stages, all originating from European stratigraphy, is now used worldwide. In many parts of the world, alternative local subdivisions are still in use.

From youngest to oldest, the subdivisions of the Cretaceous period are:

Boundaries

The lower boundary of the Cretaceous is currently undefined, and the Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined Global Boundary Stratotype Section and Point (GSSP). Placing a GSSP for this boundary has been difficult because of the strong regionality of most biostratigraphic markers, and the lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes) that could be used to define or correlate a boundary. Calpionellids, an enigmatic group of planktonic protists with urn-shaped calcitic tests briefly abundant during the latest Jurassic to earliest Cretaceous, have been suggested as the most promising candidates for fixing the Jurassic–Cretaceous boundary. In particular, the first appearance Calpionella alpina, coinciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous. The working definition for the boundary has often been placed as the first appearance of the ammonite Strambergella jacobi, formerly placed in the genus Berriasella, but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of C. alpina. The boundary is officially considered by the International Commission on Stratigraphy to be approximately 145 million years ago, but other estimates have been proposed based on U-Pb geochronology, ranging as young as 140 million years ago.

The upper boundary of the Cretaceous is sharply defined, being placed at an iridium-rich layer found worldwide that is believed to be associated with the Chicxulub impact crater, with its boundaries circumscribing parts of the Yucatán Peninsula and extending into the Gulf of Mexico. This layer has been dated at 66.043 Mya.

At the end of the Cretaceous, the impact of a large body with the Earth may have been the punctuation mark at the end of a progressive decline in biodiversity during the Maastrichtian age. The result was the extinction of three-quarters of Earth's plant and animal species. The impact created the sharp break known as the K–Pg boundary (formerly known as the K–T boundary). Earth's biodiversity required substantial time to recover from this event, despite the probable existence of an abundance of vacant ecological niches.

Despite the severity of the K-Pg extinction event, there were significant variations in the rate of extinction between and within different clades. Species that depended on photosynthesis declined or became extinct as atmospheric particles blocked solar energy. As is the case today, photosynthesizing organisms, such as phytoplankton and land plants, formed the primary part of the food chain in the late Cretaceous, and all else that depended on them suffered, as well. Herbivorous animals, which depended on plants and plankton as their food, died out as their food sources became scarce; consequently, the top predators, such as Tyrannosaurus rex, also perished. Yet only three major groups of tetrapods disappeared completely; the nonavian dinosaurs, the plesiosaurs and the pterosaurs. The other Cretaceous groups that did not survive into the Cenozoic Era—the ichthyosaurs, last remaining temnospondyls (Koolasuchus), and nonmammalian cynodonts (Tritylodontidae) — were already extinct millions of years before the event occurred.

Coccolithophorids and molluscs, including ammonites, rudists, freshwater snails, and mussels, as well as organisms whose food chain included these shell builders, became extinct or suffered heavy losses. For example, ammonites are thought to have been the principal food of mosasaurs, a group of giant marine lizards related to snakes that became extinct at the boundary.

Omnivores, insectivores, and carrion-eaters survived the extinction event, perhaps because of the increased availability of their food sources. At the end of the Cretaceous, there seem to have been no purely herbivorous or carnivorous mammals. Mammals and birds that survived the extinction fed on insects, larvae, worms, and snails, which in turn fed on dead plant and animal matter. Scientists theorise that these organisms survived the collapse of plant-based food chains because they fed on detritus.

In stream communities, few groups of animals became extinct. Stream communities rely less on food from living plants and more on detritus that washes in from land. This particular ecological niche buffered them from extinction. Similar, but more complex patterns have been found in the oceans. Extinction was more severe among animals living in the water column than among animals living on or in the seafloor. Animals in the water column are almost entirely dependent on primary production from living phytoplankton, while animals living on or in the ocean floor feed on detritus or can switch to detritus feeding.

The largest air-breathing survivors of the event, crocodilians and champsosaurs, were semiaquatic and had access to detritus. Modern crocodilians can live as scavengers and can survive for months without food and go into hibernation when conditions are unfavorable, and their young are small, grow slowly, and feed largely on invertebrates and dead organisms or fragments of organisms for their first few years. These characteristics have been linked to crocodilian survival at the end of the Cretaceous.

Geologic formations
The high sea level and warm climate of the Cretaceous meant large areas of the continents were covered by warm, shallow seas, providing habitat for many marine organisms. The Cretaceous was named for the extensive chalk deposits of this age in Europe, but in many parts of the world, the deposits from the Cretaceous are of marine limestone, a rock type that is formed under warm, shallow marine conditions. Due to the high sea level, there was extensive space for such sedimentation. Because of the relatively young age and great thickness of the system, Cretaceous rocks are evident in many areas worldwide.

Chalk is a rock type characteristic for (but not restricted to) the Cretaceous. It consists of coccoliths, microscopically small calcite skeletons of coccolithophores, a type of algae that prospered in the Cretaceous seas.

Stagnation of deep sea currents in middle Cretaceous times caused anoxic conditions in the sea water leaving the deposited organic matter undecomposed. Half of the world's petroleum reserves were laid down at this time in the anoxic conditions of what would become the Persian Gulf and the Gulf of Mexico. In many places around the world, dark anoxic shales were formed during this interval, such as the Mancos Shale of western North America. These shales are an important source rock for oil and gas, for example in the subsurface of the North Sea.

Europe
In northwestern Europe, chalk deposits from the Upper Cretaceous are characteristic for the Chalk Group, which forms the white cliffs of Dover on the south coast of England and similar cliffs on the French Normandian coast. The group is found in England, northern France, the low countries, northern Germany, Denmark and in the subsurface of the southern part of the North Sea. Chalk is not easily consolidated and the Chalk Group still consists of loose sediments in many places. The group also has other limestones and arenites. Among the fossils it contains are sea urchins, belemnites, ammonites and sea reptiles such as Mosasaurus.

In southern Europe, the Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls. Because the Alpine mountain chains did not yet exist in the Cretaceous, these deposits formed on the southern edge of the European continental shelf, at the margin of the Tethys Ocean.

North America
During the Cretaceous, the present North American continent was isolated from the other continents. In the Jurassic, the North Atlantic already opened, leaving a proto-ocean between Europe and North America. From north to south across the continent, the Western Interior Seaway started forming. This inland sea separated the elevated areas of Laramidia in the west and Appalachia in the east. Three dinosaur clades found in Laramidia (troodontids, therizinosaurids and oviraptorosaurs) are absent from Appalachia from the Coniacian through the Maastrichtian.

Paleogeography
During the Cretaceous, the late-Paleozoic-to-early-Mesozoic supercontinent of Pangaea completed its tectonic breakup into the present-day continents, although their positions were substantially different at the time. As the Atlantic Ocean widened, the convergent-margin mountain building (orogenies) that had begun during the Jurassic continued in the North American Cordillera, as the Nevadan orogeny was followed by the Sevier and Laramide orogenies.

Gondwana had begun to break up during the Jurassic Period, but its fragmentation accelerated during the Cretaceous and was largely complete by the end of the period. South America, Antarctica, and Australia rifted away from Africa (though India and Madagascar remained attached to each other until around 80 million years ago); thus, the South Atlantic and Indian Oceans were newly formed. Such active rifting lifted great undersea mountain chains along the welts, raising eustatic sea levels worldwide. To the north of Africa the Tethys Sea continued to narrow. During most of the Late Cretaceous, North America would be divided in two by the Western Interior Seaway, a large interior sea, separating Laramidia to the west and Appalachia to the east, then receded late in the period, leaving thick marine deposits sandwiched between coal beds. Bivalve palaeobiogeography also indicates that Africa was split in half by a shallow sea during the Coniacian and Santonian, connecting the Tethys with the South Atlantic by way of the central Sahara and Central Africa, which were then underwater. Yet another shallow seaway ran between what is now Norway and Greenland, connecting the Tethys to the Arctic Ocean and enabling biotic exchange between the two oceans. At the peak of the Cretaceous transgression, one-third of Earth's present land area was submerged.

The Cretaceous is justly famous for its chalk; indeed, more chalk formed in the Cretaceous than in any other period in the Phanerozoic. Mid-ocean ridge activity—or rather, the circulation of seawater through the enlarged ridges—enriched the oceans in calcium; this made the oceans more saturated, as well as increased the bioavailability of the element for calcareous nanoplankton. These widespread carbonates and other sedimentary deposits make the Cretaceous rock record especially fine. Famous formations from North America include the rich marine fossils of Kansas's Smoky Hill Chalk Member and the terrestrial fauna of the late Cretaceous Hell Creek Formation. Other important Cretaceous exposures occur in Europe (e.g., the Weald) and China (the Yixian Formation). In the area that is now India, massive lava beds called the Deccan Traps were erupted in the very late Cretaceous and early Paleocene.

Climate
Palynological evidence indicates the Cretaceous climate had three broad phases: a Berriasian–Barremian warm-dry phase, an Aptian–Santonian warm-wet phase, and a Campanian–Maastrichtian cool-dry phase. As in the Cenozoic, the 400,000 year eccentricity cycle was the dominant orbital cycle governing carbon flux between different reservoirs and influencing global climate. The location of the Intertropical Convergence Zone (ITCZ) was roughly the same as in the present.

The cooling trend of the last epoch of the Jurassic, the Tithonian, continued into the Berriasian, the first age of the Cretaceous. The North Atlantic seaway opened and enabled the flow of cool water from the Boreal Ocean into the Tethys. There is evidence that snowfalls were common in the higher latitudes during this age, and the tropics became wetter than during the Triassic and Jurassic. Glaciation was restricted to high-latitude mountains, though seasonal snow may have existed farther from the poles. After the end of the first age, however, temperatures began to increase again, with a number of thermal excursions, such as the middle Valanginian Weissert Thermal Excursion (WTX), which was caused by the Paraná-Etendeka Large Igneous Province's activity. It was followed by the middle Hauterivian Faraoni Thermal Excursion (FTX) and the early Barremian Hauptblatterton Thermal Event (HTE). The HTE marked the ultimate end of the Tithonian-early Barremian Cool Interval (TEBCI). During this interval, precession was the dominant orbital driver of environmental changes in the Vocontian Basin. For much of the TEBCI, northern Gondwana experienced a monsoonal climate. A shallow thermocline existed in the mid-latitude Tethys. The TEBCI was followed by the Barremian-Aptian Warm Interval (BAWI). This hot climatic interval coincides with Manihiki and Ontong Java Plateau volcanism and with the Selli Event. Early Aptian tropical sea surface temperatures (SSTs) were 27–32 °C, based on TEX86 measurements from the equatorial Pacific. During the Aptian, Milankovitch cycles governed the occurrence of anoxic events by modulating the intensity of the hydrological cycle and terrestrial runoff. The early Aptian was also notable for its millennial scale hyperarid events in the mid-latitudes of Asia. The BAWI itself was followed by the Aptian-Albian Cold Snap (AACS) that began about 118 Ma. A short, relatively minor ice age may have occurred during this so-called "cold snap", as evidenced by glacial dropstones in the western parts of the Tethys Ocean and the expansion of calcareous nannofossils that dwelt in cold water into lower latitudes. The AACS is associated with an arid period in the Iberian Peninsula.

Temperatures increased drastically after the end of the AACS, which ended around 111 Ma with the Paquier/Urbino Thermal Maximum, giving way to the Mid-Cretaceous Hothouse (MKH), which lasted from the early Albian until the early Campanian. Faster rates of seafloor spreading and entry of carbon dioxide into the atmosphere are believed to have initiated this period of extreme warmth, along with high flood basalt activity. The MKH was punctuated by multiple thermal maxima of extreme warmth. The Leenhardt Thermal Event (LTE) occurred around 110 Ma, followed shortly by the l’Arboudeyesse Thermal Event (ATE) a million years later. Following these two hyperthermals was the Amadeus Thermal Maximum around 106 Ma, during the middle Albian. Then, around a million years after that, occurred the Petite Verol Thermal Event (PVTE). Afterwards, around 102.5 Ma, the Event 6 Thermal Event (EV6) took place; this event was itself followed by the Breistroffer Thermal Maximum around 101 Ma, during the latest Albian. Approximately 94 Ma, the Cenomanian-Turonian Thermal Maximum occurred, with this hyperthermal being the most extreme hothouse interval of the Cretaceous  and being associated with a sea level highstand. Temperatures cooled down slightly over the next few million years, but then another thermal maximum, the Coniacian Thermal Maximum, happened, with this thermal event being dated to around 87 Ma. Atmospheric CO2 levels may have varied by thousands of ppm throughout the MKH. Mean annual temperatures at the poles during the MKH exceeded 14 °C. Such hot temperatures during the MKH resulted in a very gentle temperature gradient from the equator to the poles; the latitudinal temperature gradient during the Cenomanian-Turonian Thermal Maximum was 0.54 °C per ° latitude for the Southern Hemisphere and 0.49 °C per ° latitude for the Northern Hemisphere, in contrast to present day values of 1.07 and 0.69 °C per ° latitude for the Southern and Northern hemispheres, respectively. This meant weaker global winds, which drive the ocean currents, and resulted in less upwelling and more stagnant oceans than today. This is evidenced by widespread black shale deposition and frequent anoxic events. Tropical SSTs during the late Albian most likely averaged around 30 °C. Despite this high SST, seawater was not hypersaline at this time, as this would have required significantly higher temperatures still. On land, arid zones in the Albian regularly expanded northward in tandem with expansions of subtropical high pressure belts. The Cedar Mountain Formation's Soap Wash flora indicates a mean annual temperature of between 19-26 °C in Utah at the Albian-Cenomanian boundary. Tropical SSTs during the Cenomanian-Turonian Thermal Maximum were at least 30 °C, though one study estimated them as high as between 33 and 42 °C. An intermediate estimate of ~33-34 °C has also been given. Meanwhile, deep ocean temperatures were as much as 15 to 20 C-change warmer than today's; one study estimated that deep ocean temperatures were between 12 and 20 °C during the MKH. The poles were so warm that ectothermic reptiles were able to inhabit them.

Beginning in the Santonian, near the end of the MKH, the global climate began to cool, with this cooling trend continuing across the Campanian. This period of cooling, driven by falling levels of atmospheric carbon dioxide, caused the end of the MKH and the transition into a cooler climatic interval, known formally as the Late Cretaceous-Early Palaeogene Cool Interval (LKEPCI). Tropical SSTs declined from around 35 °C in the early Campanian to around 28 °C in the Maastrichtian. Deep ocean temperatures declined to 9 to 12 °C, though the shallow temperature gradient between tropical and polar seas remained. Regional conditions in the Western Interior Seaway changed little between the MKH and the LKEPCI. During this period of relatively cool temperatures, the ITCZ became narrower, while the strength of both summer and winter monsoons in East Asia was directly correlated to atmospheric CO2 concentrations. Laramidia likewise had a seasonal, monsoonal climate. The Maastrichtian was a time of chaotic, highly variable climate. Two upticks in global temperatures are known to have occurred during the Maastrichtian, bucking the trend of overall cooler temperatures during the LKEPCI. Between 70 and 69 Ma and 66–65 Ma, isotopic ratios indicate elevated atmospheric CO2 pressures with levels of 1000–1400 ppmV and mean annual temperatures in west Texas between 21 and 23 C. Atmospheric CO2 and temperature relations indicate a doubling of pCO2 was accompanied by a ~0.6 °C increase in temperature. The latter warming interval, occurring at the very end of the Cretaceous, was triggered by the activity of the Deccan Traps. The LKEPCI lasted into the Late Palaeocene, when it gave way to another supergreenhouse interval.

The production of large quantities of magma, variously attributed to mantle plumes or to extensional tectonics, further pushed sea levels up, so that large areas of the continental crust were covered with shallow seas. The Tethys Sea connecting the tropical oceans east to west also helped to warm the global climate. Warm-adapted plant fossils are known from localities as far north as Alaska and Greenland, while dinosaur fossils have been found within 15 degrees of the Cretaceous south pole. It was suggested that there was Antarctic marine glaciation in the Turonian Age, based on isotopic evidence. However, this has subsequently been suggested to be the result of inconsistent isotopic proxies, with evidence of polar rainforests during this time interval at 82° S. Rafting by ice of stones into marine environments occurred during much of the Cretaceous, but evidence of deposition directly from glaciers is limited to the Early Cretaceous of the Eromanga Basin in southern Australia.

Flora

Flowering plants (angiosperms) make up around 90% of living plant species today. Prior to the rise of angiosperms, during the Jurassic and the Early Cretaceous, the higher flora was dominated by gymnosperm groups, including cycads, conifers, ginkgophytes, gnetophytes and close relatives, as well as the extinct Bennettitales. Other groups of plants included pteridosperms or "seed ferns", a collective term that refers to disparate groups of extinct seed plants with fern-like foliage, including groups such as Corystospermaceae and Caytoniales. The exact origins of angiosperms are uncertain, although molecular evidence suggests that they are not closely related to any living group of gymnosperms.

The earliest widely accepted evidence of flowering plants are monosulcate (single-grooved) pollen grains from the late Valanginian (~ 134 million years ago) found in Israel and Italy, initially at low abundance. Molecular clock estimates conflict with fossil estimates, suggesting the diversification of crown-group angiosperms during the Late Triassic or the Jurassic, but such estimates are difficult to reconcile with the heavily sampled pollen record and the distinctive tricolpate to tricolporoidate (triple grooved) pollen of eudicot angiosperms. Among the oldest records of Angiosperm macrofossils are Montsechia from the Barremian aged Las Hoyas beds of Spain and Archaefructus from the Barremian-Aptian boundary Yixian Formation in China. Tricolpate pollen distinctive of eudicots first appears in the Late Barremian, while the earliest remains of monocots are known from the Aptian. Flowering plants underwent a rapid radiation beginning during the middle Cretaceous, becoming the dominant group of land plants by the end of the period, coincident with the decline of previously dominant groups such as conifers. The oldest known fossils of grasses are from the Albian, with the family having diversified into modern groups by the end of the Cretaceous. The oldest large angiosperm trees are known from the Turonian (c. 90 Mya) of New Jersey, with the trunk having a preserved diameter of 1.8 m and an estimated height of 50 m.

During the Cretaceous, ferns in the order Polypodiales, which make up 80% of living fern species, would also begin to diversify.

Terrestrial fauna
On land, mammals were generally small sized, but a very relevant component of the fauna, with cimolodont multituberculates outnumbering dinosaurs in some sites. Neither true marsupials nor placentals existed until the very end, but a variety of non-marsupial metatherians and non-placental eutherians had already begun to diversify greatly, ranging as carnivores (Deltatheroida), aquatic foragers (Stagodontidae) and herbivores (Schowalteria, Zhelestidae). Various "archaic" groups like eutriconodonts were common in the Early Cretaceous, but by the Late Cretaceous northern mammalian faunas were dominated by multituberculates and therians, with dryolestoids dominating South America.

The apex predators were archosaurian reptiles, especially dinosaurs, which were at their most diverse stage. Avians such as the ancestors of modern-day birds also diversified. They inhabited every continent, and were even found in cold polar latitudes. Pterosaurs were common in the early and middle Cretaceous, but as the Cretaceous proceeded they declined for poorly understood reasons (once thought to be due to competition with early birds, but now it is understood avian adaptive radiation is not consistent with pterosaur decline ). By the end of the period only three highly specialized families remained; Pteranodontidae, Nyctosauridae, and Azhdarchidae.

The Liaoning lagerstätte (Yixian Formation) in China is an important site, full of preserved remains of numerous types of small dinosaurs, birds and mammals, that provides a glimpse of life in the Early Cretaceous. The coelurosaur dinosaurs found there represent types of the group Maniraptora, which includes modern birds and their closest non-avian relatives, such as dromaeosaurs, oviraptorosaurs, therizinosaurs, troodontids along with other avialans. Fossils of these dinosaurs from the Liaoning lagerstätte are notable for the presence of hair-like feathers.

Insects diversified during the Cretaceous, and the oldest known ants, termites and some lepidopterans, akin to butterflies and moths, appeared. Aphids, grasshoppers and gall wasps appeared.

Rhynchocephalians
Rhynchocephalians (which today only includes the tuatara) disappeared from North America and Europe after the Early Cretaceous, and were absent from North Africa and northern South America by the early Late Cretaceous. The cause of the decline of Rhynchocephalia remains unclear, but has often been suggested to be due to competition with advanced lizards and mammals. They appear to have remained diverse in high-latitude southern South America during the Late Cretaceous, where lizards remained rare, with their remains outnumbering terrestrial lizards 200:1.

Choristodera
Choristoderes, a group of freshwater aquatic reptiles that first appeared during the preceding Jurassic, underwent a major evolutionary radiation in Asia during the Early Cretaceous, which represents the high point of choristoderan diversity, including long necked forms such as Hyphalosaurus and the first records of the gharial-like Neochoristodera, which appear to have evolved in the regional absence of aquatic neosuchian crocodyliformes. During the Late Cretaceous the neochoristodere Champsosaurus was widely distributed across western North America. Due to the extreme climatic warmth in the Arctic, choristoderans were able to colonise it too during the Late Cretaceous.

Marine fauna
In the seas, rays, modern sharks and teleosts became common. Marine reptiles included ichthyosaurs in the early and mid-Cretaceous (becoming extinct during the late Cretaceous Cenomanian-Turonian anoxic event), plesiosaurs throughout the entire period, and mosasaurs appearing in the Late Cretaceous. Sea turtles in the form of Cheloniidae and Panchelonioidea lived during the period and survived the extinction event. Panchelonioidea is today represented by a single species; the leatherback sea turtle. The Hesperornithiformes were flightless, marine diving birds that swam like grebes.

Baculites, an ammonite genus with a straight shell, flourished in the seas along with reef-building rudist clams. Inoceramids were also particularly notable among Cretaceous bivalves, and they have been used to identify major biotic turnovers such as at the Turonian-Coniacian boundary. Predatory gastropods with drilling habits were widespread. Globotruncanid foraminifera and echinoderms such as sea urchins and starfish (sea stars) thrived. Ostracods were abundant in Cretaceous marine settings; ostracod species characterised by high male sexual investment had the highest rates of extinction and turnover. Thylacocephala, a class of crustaceans, went extinct in the Late Cretaceous. The first radiation of the diatoms (generally siliceous shelled, rather than calcareous) in the oceans occurred during the Cretaceous; freshwater diatoms did not appear until the Miocene. Calcareous nannoplankton were important components of the marine microbiota and important as biostratigraphic markers and recorders of environmental change.

The Cretaceous was also an important interval in the evolution of bioerosion, the production of borings and scrapings in rocks, hardgrounds and shells.

title: Cretaceous


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Creutzfeldt–Jakob disease

Creutzfeldt–Jakob disease (CJD), also known as subacute spongiform encephalopathy or neurocognitive disorder due to prion disease, is a fatal neurodegenerative disease. Early symptoms include memory problems, behavioral changes, poor coordination, and visual disturbances. Later symptoms include dementia, involuntary movements, blindness, weakness, and coma. About 70% of people die within a year of diagnosis. The name "Creutzfeldt–Jakob disease" was introduced by Walther Spielmeyer in 1922, after the German neurologists Hans Gerhard Creutzfeldt and Alfons Maria Jakob.

CJD is caused by a type of abnormal shaping of a protein known as a prion. Infectious prions are misfolded proteins that can cause normally folded proteins to also become misfolded. About 85% of cases of CJD occur for unknown reasons, while about 7.5% of cases are inherited in an autosomal dominant manner. Exposure to brain or spinal tissue from an infected person may also result in spread. There is no evidence that sporadic CJD can spread among people via normal contact or blood transfusions, although this is possible in variant Creutzfeldt–Jakob disease. Diagnosis involves ruling out other potential causes. An electroencephalogram, spinal tap, or magnetic resonance imaging may support the diagnosis.

There is no specific treatment for CJD. Opioids may be used to help with pain, while clonazepam or sodium valproate may help with involuntary movements. CJD affects about one person per million people per year. Onset is typically around 60 years of age. The condition was first described in 1920. It is classified as a type of transmissible spongiform encephalopathy. Inherited CJD accounts for about 10% of prion disease cases. Sporadic CJD is different from bovine spongiform encephalopathy (mad cow disease) and variant Creutzfeldt–Jakob disease (vCJD).

Signs and symptoms
The first symptom of CJD is usually rapidly progressive dementia, leading to memory loss, personality changes, and hallucinations. Myoclonus (jerky movements) typically occurs in 90% of cases, but may be absent at initial onset. Other frequently occurring features include anxiety, depression, paranoia, obsessive-compulsive symptoms, and psychosis. This is accompanied by physical problems such as speech impairment, balance and coordination dysfunction (ataxia), changes in gait, and rigid posture. In most people with CJD, these symptoms are accompanied by involuntary movements. The duration of the disease varies greatly, but sporadic (non-inherited) CJD can be fatal within months or even weeks. Most affected people die six months after initial symptoms appear, often of pneumonia due to impaired coughing reflexes. About 15% of people with CJD survive for two or more years.

The symptoms of CJD are caused by the progressive death of the brain's nerve cells, which are associated with the build-up of abnormal prion proteins forming in the brain. When brain tissue from a person with CJD is examined under a microscope, many tiny holes can be seen where the nerve cells have died. Parts of the brain may resemble a sponge where the prions were infecting the areas of the brain.

Cause
CJD is a type of transmissible spongiform encephalopathy (TSE), which are caused by prions. Prions are misfolded proteins that occur in the neurons of the central nervous system (CNS). They are thought to affect signaling processes, damaging neurons and resulting in degeneration that causes the spongiform appearance in the affected brain.

The CJD prion is dangerous because it promotes refolding of the cellular prion protein into the diseased state. The number of misfolded protein molecules will increase exponentially and the process leads to a large quantity of insoluble protein in affected cells. This mass of misfolded proteins disrupts neuronal cell function and causes cell death. Mutations in the gene for the prion protein can cause a misfolding of the dominantly alpha helical regions into beta pleated sheets. This change in conformation disables the ability of the protein to undergo digestion. Once the prion is transmitted, the defective proteins invade the brain and induce other prion protein molecules to misfold in a self-sustaining feedback loop. These neurodegenerative diseases are commonly called prion diseases.

Transmission
The defective protein can be transmitted by contaminated harvested human brain products, corneal grafts, dural grafts, or electrode implants and human growth hormone.

It can be familial (fCJD); or it may appear without clear risk factors (sporadic form: sCJD). In the familial form, a mutation has occurred in the gene for PrP, PRNP, in that family. All types of CJD are transmissible irrespective of how they occur in the person.

It is thought that humans can contract the variant form of the disease by eating food from animals infected with bovine spongiform encephalopathy (BSE), the bovine form of TSE also known as mad cow disease. However, it can also cause sCJD in some cases.

Cannibalism has also been implicated as a transmission mechanism for abnormal prions, causing the disease known as kuru, once found primarily among women and children of the Fore people in Papua New Guinea, who previously engaged in funerary cannibalism. While the men of the tribe ate the muscle tissue of the deceased, women and children consumed other parts, such as the brain, and were more likely than men to contract kuru from infected tissue.

Prions, the infectious agent of CJD, may not be inactivated by means of routine surgical instrument sterilization procedures. The World Health Organization and the US Centers for Disease Control and Prevention recommend that instrumentation used in such cases be immediately destroyed after use; short of destruction, it is recommended that heat and chemical decontamination be used in combination to process instruments that come in contact with high-infectivity tissues. Thermal depolymerization also destroys prions in infected organic and inorganic matter, since the process chemically attacks protein at the molecular level, although more effective and practical methods involve destruction by combinations of detergents and enzymes similar to biological washing powders.

Genetics
People can also develop CJD because they carry a mutation of the gene that codes for the prion protein (PRNP), located on chromosome 202p12-pter. This occurs in only 10–15% of all CJD cases. In sporadic cases, the misfolding of the prion protein is a process that is hypothesized to occur as a result of the effects of aging on cellular machinery, explaining why the disease often appears later in life. An EU study determined that "87% of cases were sporadic, 8% genetic, 5% iatrogenic and less than 1% variant."

Diagnosis
Testing for CJD has historically been problematic, due to nonspecific nature of early symptoms and difficulty in safely obtaining brain tissue for confirmation. The diagnosis may initially be suspected in a person with rapidly progressing dementia, particularly when they are also found with the characteristic medical signs and symptoms such as involuntary muscle jerking, difficulty with coordination/balance and walking, and visual disturbances. Further testing can support the diagnosis and may include:
 * Electroencephalography – may have characteristic generalized periodic sharp wave pattern. Periodic sharp wave complexes develop in half of the people with sporadic CJD, particularly in the later stages.
 * Cerebrospinal fluid (CSF) analysis for elevated levels of 14-3-3 protein could be supportive in the diagnosis of sCJD. However, a positive result should not be regarded as sufficient for the diagnosis.  The Real-Time Quaking-Induced Conversion (RT-QuIC) assay has a diagnostic sensitivity of more than 80% and a specificity approaching 100%, tested in detecting PrPSc in CSF samples of people with CJD.   It is therefore suggested as a high-value diagnostic method for the disease.
 * MRI of the brain – often shows high signal intensity in the caudate nucleus and putamen bilaterally on T2-weighted images.

In recent years, studies have shown that the tumour marker neuron-specific enolase (NSE) is often elevated in CJD cases; however, its diagnostic utility is seen primarily when combined with a test for the 14-3-3 protein. , screening tests to identify infected asymptomatic individuals, such as blood donors, are not yet available, though methods have been proposed and evaluated.

Imaging
Imaging of the brain may be performed during medical evaluation, both to rule out other causes and to obtain supportive evidence for diagnosis. Imaging findings are variable in their appearance, and also variable in sensitivity and specificity. While imaging plays a lesser role in diagnosis of CJD, characteristic findings on brain MRI in some cases may precede onset of clinical manifestations.

Brain MRI is the most useful imaging modality for changes related to CJD. Of the MRI sequences, diffuse-weighted imaging sequences are most sensitive. Characteristic findings are as follows:
 * Focal or diffuse diffusion-restriction involving the cerebral cortex and/or basal ganglia. In about 24% of cases DWI shows only cortical hyperintensity; in 68%, cortical and subcortical abnormalities; and in 5%, only subcortical anomalies. The most iconic and striking cortical abnormality has been called "cortical ribboning" or "cortical ribbon sign" due to hyperintensities resembling ribbons appearing in the cortex on MRI. The involvement of the thalamus can be found in sCJD, is even stronger and constant in vCJD.
 * Varying degree of symmetric T2 hyperintense signal changes in the basal ganglia (i.e., caudate and putamen), and to a lesser extent globus pallidus and occipital cortex.
 * Cerebellar atrophy

Brain FDG PET-CT tends to be markedly abnormal, and is increasingly used in the investigation of dementias.
 * Patients with CJD will normally have hypometabolism on FDG PET.

Histopathology
Testing of tissue remains the most definitive way of confirming the diagnosis of CJD, although it must be recognized that even biopsy is not always conclusive.

In one-third of people with sporadic CJD, deposits of "prion protein (scrapie)", PrPSc, can be found in the skeletal muscle and/or the spleen. Diagnosis of vCJD can be supported by biopsy of the tonsils, which harbor significant amounts of PrPSc; however, biopsy of brain tissue is the definitive diagnostic test for all other forms of prion disease. Due to its invasiveness, biopsy will not be done if clinical suspicion is sufficiently high or low. A negative biopsy does not rule out CJD, since it may predominate in a specific part of the brain.

The classic histologic appearance is spongiform change in the gray matter: the presence of many round vacuoles from one to 50 micrometers in the neuropil, in all six cortical layers in the cerebral cortex or with diffuse involvement of the cerebellar molecular layer. These vacuoles appear glassy or eosinophilic and may coalesce. Neuronal loss and gliosis are also seen. Plaques of amyloid-like material can be seen in the neocortex in some cases of CJD.

However, extra-neuronal vacuolization can also be seen in other disease states. Diffuse cortical vacuolization occurs in Alzheimer's disease, and superficial cortical vacuolization occurs in ischemia and frontotemporal dementia. These vacuoles appear clear and punched-out. Larger vacuoles encircling neurons, vessels, and glia are a possible processing artifact.

Classification
Types of CJD include:
 * Sporadic (sCJD), caused by the spontaneous misfolding of prion-protein in an individual. This accounts for 85% of cases of CJD.
 * Familial (fCJD), caused by an inherited mutation in the prion-protein gene. This accounts for the majority of the other 15% of cases of CJD.
 * Acquired CJD, caused by contamination with tissue from an infected person, usually as the result of a medical procedure (iatrogenic CJD). Medical procedures that are associated with the spread of this form of CJD include blood transfusion from the infected person, use of human-derived pituitary growth hormones, gonadotropin hormone therapy, and corneal and meningeal transplants. Variant Creutzfeldt–Jakob disease (vCJD) is a type of acquired CJD potentially acquired from bovine spongiform encephalopathy or caused by consuming food contaminated with prions.

Treatment
As of 2024, there is no cure or effective treatment for CJD. Some of the symptoms like twitching can be managed, but otherwise treatment is palliative care. Psychiatric symptoms like anxiety and depression can be treated with sedatives and antidepressants. Myoclonic jerks can be handled with clonazepam or sodium valproate. Opiates can help in pain. Seizures are very uncommon but can nevertheless be treated with antiepileptic drugs.

Prognosis
Life expectancy is greatly reduced for people with Creutzfeldt-Jakob disease, with the average being less than 6 months. As of 1981, no one was known to have lived longer than 2.5 years after the onset of CJD symptoms. In 2011, Jonathan Simms, a Northern Irish man who lived 10 years after his diagnosis, was reported to be one of the world's longest survivors of variant Creutzfeldt–Jakob disease (vCJD).

Epidemiology
CDC monitors the occurrence of CJD in the United States through periodic reviews of national mortality data. According to the CDC:
 * CJD occurs worldwide at a rate of about 1 case per million population per year.
 * On the basis of mortality surveillance from 1979 to 1994, the annual incidence of CJD remained stable at approximately 1 case per million people in the United States.
 * In the United States, CJD deaths among people younger than 30 years of age are extremely rare (fewer than five deaths per billion per year).
 * The disease is found most frequently in people 55–65 years of age, but cases can occur in people older than 90 years and younger than 55 years of age.
 * In more than 85% of cases, the duration of CJD is less than one year (median: four months) after the onset of symptoms.

Further information from the CDC:
 * Risk of developing CJD increases with age.
 * CJD incidence was 3.5 cases per million among those over 50 years of age between 1979 and 2017.
 * Approximately 85% of CJD cases are sporadic and 10-15% of CJD cases are due to inherited mutations of the prion protein gene.
 * CJD deaths and age-adjusted death rate in the United States indicate an increasing trend in the number of deaths between 1979 and 2017.

Although not fully understood, additional information suggests that CJD rates in African American and nonwhite groups are lower than in whites. While the mean onset is approximately 67 years of age, cases of sCJD have been reported as young as 17 years and over 80 years of age. Mental capabilities rapidly deteriorate and the average amount of time from onset of symptoms to death is 7 to 9 months.

According to a 2020 systematic review on the international epidemiology of CJD:
 * Surveillance studies from 2005 and later show the estimated global incidence is 1–2 cases per million population per year.
 * Sporadic CJD (sCJD) incidence increased from the years 1990–2018 in the UK.
 * Probable or definite sCJD deaths also increased from the years 1996–2018 in twelve additional countries.
 * CJD incidence is greatest in those over the age of 55 years old, with an average age of 67 years old.

The intensity of CJD surveillance increases the number of reported cases, often in countries where CJD epidemics have occurred in the past and where surveillance resources are greatest. An increase in surveillance and reporting of CJD is most likely in response to BSE and vCJD. Possible factors contributing to an increase of CJD incidence are an aging population, population increase, clinician awareness, and more accurate diagnostic methods. Since CJD symptoms are similar to other neurological conditions, it is also possible that CJD is mistaken for stroke, acute nephropathy, general dementia, and hyperparathyroidism.

History
The disease was first described by German neurologist Hans Gerhard Creutzfeldt in 1920 and shortly afterward by Alfons Maria Jakob, giving it the name Creutzfeldt–Jakob disease. Some of the clinical findings described in their first papers do not match current criteria for Creutzfeldt–Jakob disease, and it has been speculated that at least two of the people in initial studies had a different ailment. An early description of familial CJD stems from the German psychiatrist and neurologist Friedrich Meggendorfer (1880–1953). A study published in 1997 counted more than 100 cases worldwide of transmissible CJD and new cases continued to appear at the time.

The first report of suspected iatrogenic CJD was published in 1974. Animal experiments showed that corneas of infected animals could transmit CJD, and the causative agent spreads along visual pathways. A second case of CJD associated with a corneal transplant was reported without details. In 1977, CJD transmission caused by silver electrodes previously used in the brain of a person with CJD was first reported. Transmission occurred despite the decontamination of the electrodes with ethanol and formaldehyde. Retrospective studies identified four other cases likely of similar cause. The rate of transmission from a single contaminated instrument is unknown, although it is not 100%. In some cases, the exposure occurred weeks after the instruments were used on a person with CJD. In the 1980s it was discovered that Lyodura, a dura mater transplant product, was shown to transmit CJD from the donor to the recipient. This led to the product being banned in Canada but it was used in other countries such as Japan until 1993.

A review article published in 1979 indicated that 25 dura mater cases had occurred by that date in Australia, Canada, Germany, Italy, Japan, New Zealand, Spain, the United Kingdom, and the United States.

By 1985, a series of case reports in the United States showed that when injected, cadaver-extracted pituitary human growth hormone could transmit CJD to humans.

In 1992, it was recognized that human gonadotropin administered by injection could also transmit CJD from person to person.

Stanley B. Prusiner of the University of California, San Francisco (UCSF) was awarded the Nobel Prize in Physiology or Medicine in 1997 "for his discovery of Prions—a new biological principle of infection".

Yale University neuropathologist Laura Manuelidis has challenged the prion protein (PrP) explanation for the disease. In January 2007, she and her colleagues reported that they had found a virus-like particle in naturally and experimentally infected animals. "The high infectivity of comparable, isolated virus-like particles that show no intrinsic PrP by antibody labeling, combined with their loss of infectivity when nucleic acid–protein complexes are disrupted, make it likely that these 25-nm particles are the causal TSE virions".

Australia
Australia has documented 10 cases of healthcare-acquired CJD (iatrogenic or ICJD). Five of the deaths resulted after the patients, who were in treatment either for infertility or short stature, were treated using contaminated pituitary extract hormone but no new cases have been noted since 1991. The other five deaths occurred due to dura grafting procedures that were performed during brain surgery, in which the covering of the brain is repaired. There have been no other ICJD deaths documented in Australia due to transmission during healthcare procedures.

New Zealand
A case was reported in 1989 in a 25-year-old man from New Zealand, who also received dura mater transplant. Five New Zealanders have been confirmed to have died of the sporadic form of Creutzfeldt–Jakob disease (CJD) in 2012.

United States
In 1988, there was a confirmed death from CJD of a person from Manchester, New Hampshire. Massachusetts General Hospital believed the person acquired the disease from a surgical instrument at a podiatrist's office. In 2007, Michael Homer, former Vice President of Netscape, had been experiencing consistent memory problems which led to his diagnosis. In September 2013, another person in Manchester was posthumously determined to have died of the disease. The person had undergone brain surgery at Catholic Medical Center three months before his death, and a surgical probe used in the procedure was subsequently reused in other operations. Public health officials identified thirteen people at three hospitals who may have been exposed to the disease through the contaminated probe, but said the risk of anyone's contracting CJD is "extremely low". In January 2015, former speaker of the Utah House of Representatives Rebecca D. Lockhart died of the disease within a few weeks of diagnosis. John Carroll, former editor of The Baltimore Sun and Los Angeles Times, died of CJD in Kentucky in June 2015, after having been diagnosed in January. American actress Barbara Tarbuck (General Hospital, American Horror Story) died of the disease on December 26, 2016. José Baselga, clinical oncologist having headed the AstraZeneca Oncology division, died in Cerdanya, March 21, 2021, from CJD. In April 2024, two hunters from the same lodge were found to be afflicted with Sporadic CJD after eating deer meat infected with chronic wasting disease (CWD), suggesting a potential link between CWD and CJD.

Diagnosis

 * In 2010, a team from New York described detection of PrPSc in sheep's blood, even when initially present at only one part in one hundred billion (10−11) in sheep's brain tissue. The method combines amplification with a novel technology called surround optical fiber immunoassay (SOFIA) and some specific antibodies against PrPSc. The technique allowed improved detection and testing time for PrPSc.
 * In 2014, a human study showed a nasal brushing method that can accurately detect PrP in the olfactory epithelial cells of people with CJD.

Treatment

 * Pentosan polysulphate (PPS) may slow the progression of the disease, and may have contributed to the longer than expected survival of the seven people studied. The CJD Therapy Advisory Group to the UK Health Departments advises that data are not sufficient to support claims that pentosan polysulphate is an effective treatment and suggests that further research in animal models is appropriate. A 2007 review of the treatment of 26 people with PPS finds no proof of efficacy because of the lack of accepted objective criteria, but it was unclear to the authors whether that was caused by PPS itself. In 2012 it was claimed that the lack of significant benefits has likely been caused because of the drug being administered very late in the disease in many patients.
 * Use of RNA interference to slow the progression of scrapie has been studied in mice. The RNA blocks production of the protein that the CJD process transforms into prions.
 * Both amphotericin B and doxorubicin have been investigated as treatments for CJD, but as yet there is no strong evidence that either drug is effective in stopping the disease. Further study has been taken with other medical drugs, but none are effective. However, anticonvulsants and anxiolytic agents, such as valproate or a benzodiazepine, may be administered to relieve associated symptoms.
 * Quinacrine, a medicine originally created for malaria, has been evaluated as a treatment for CJD. The efficacy of quinacrine was assessed in a rigorous clinical trial in the UK and the results were published in Lancet Neurology, and concluded that quinacrine had no measurable effect on the clinical course of CJD.
 * Astemizole, a medication approved for human use, has been found to have anti-prion activity and may lead to a treatment for Creutzfeldt–Jakob disease.
 * A monoclonal antibody (code name PRN100) targeting the prion protein (PrP) was given to six people with Creutzfeldt–Jakob disease in an early-stage clinical trial conducted from 2018 to 2022. The treatment appeared to be well-tolerated and was able to access the brain, where it might have helped to clear PrPC. While the treated patients still showed progressive neurological decline, and while none of them survived longer than expected from the normal course of the disease, the scientists at University College London who conducted the study see these early-stage results as encouraging and suggest to conduct a larger study, ideally at the earliest possible intervention.

title: Creutzfeldt–Jakob disease