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Molière

Jean-Baptiste Poquelin (15 January 1622 (baptised) – 17 February 1673), known by his stage name Molière, was a French playwright, actor, and poet, widely regarded as one of the great writers in the French language and world literature. His extant works include comedies, farces, tragicomedies, comédie-ballets, and more. His plays have been translated into every major living language and are performed at the Comédie-Française more often than those of any other playwright today. His influence is such that the French language is often referred to as the "language of Molière".

Born into a prosperous family and having studied at the Collège de Clermont (now Lycée Louis-le-Grand), Molière was well suited to begin a life in the theatre. Thirteen years as an itinerant actor helped him polish his comedic abilities while he began writing, combining Commedia dell'arte elements with the more refined French comedy.

Through the patronage of aristocrats including Philippe I, Duke of Orléans—the brother of Louis XIV—Molière procured a command performance before the King at the Louvre. Performing a classic play by Pierre Corneille and a farce of his own, The Doctor in Love, Molière was granted the use of salle du Petit-Bourbon near the Louvre, a spacious room appointed for theatrical performances. Later, he was granted the use of the theatre in the Palais-Royal. In both locations, Molière found success among Parisians with plays such as The Affected Ladies, The School for Husbands, and The School for Wives. This royal favour brought a royal pension to his troupe and the title Troupe du Roi ("The King's Troupe"). Molière continued as the official author of court entertainments.

Despite the adulation of the court and Parisians, Molière's satires attracted criticism from other circles. For Tartuffe's impiety, the Catholic Church in France denounced this study of religious hypocrisy, which was followed by a ban by the Parlement, while Dom Juan was withdrawn and never restaged by Molière. His hard work in so many theatrical capacities took its toll on his health and, by 1667, he was forced to take a break from the stage. In 1673, during a production of his final play, The Imaginary Invalid, Molière, who suffered from pulmonary tuberculosis, was seized by a coughing fit and a haemorrhage while playing the hypochondriac Argan; he finished the performance but collapsed again and died a few hours later.

Life

Molière was born in Paris shortly before his christening as Jean Poquelin on 15 January 1622. Known as Jean-Baptiste, he was the first son of Jean Poquelin and Marie Cressé, who had married on 27 April 1621. His mother was the daughter of a prosperous bourgeois family. Upon seeing him for the first time, a maid exclaimed, "Le nez!", a reference to the infant's large nose. Molière was called "Le Nez" by his family from that time. He lost his mother when he was 10, and he does not seem to have been particularly close to his father. After his mother's death, he lived with his father above the Pavillon des Singes on the rue Saint-Honoré, an affluent area of Paris. It is likely that his education commenced with studies at a Parisian elementary school, followed by his enrollment in the prestigious Jesuit Collège de Clermont, where he completed his studies in a strict academic environment and got a first taste of life on the stage.

In 1631, his father Jean Poquelin purchased from the court of Louis XIII the posts of "" ("valet of the King's chamber and keeper of carpets and upholstery"). His son assumed the same posts in 1641. The title required only three months' work and an initial cost of 1,200 livres; the title paid 300 livres a year and provided a number of lucrative contracts. Molière also studied as a provincial lawyer some time around 1642, probably in Orléans, but it is not documented that he ever qualified. So far he had followed his father's plans, which had served him well; he had mingled with nobility at the Collège de Clermont and seemed destined for a career in office.

In June 1643, when Molière was 21, he decided to abandon his social class and pursue a career on the stage. Taking leave of his father, he joined the actress Madeleine Béjart, with whom he had crossed paths before, and founded the Illustre Théâtre with 630 livres. They were later joined by Madeleine's brother and sister.

The theatre troupe went bankrupt in 1645. Molière had become head of the troupe, due in part, perhaps, to his acting prowess and his legal training. However, the troupe had acquired large debts, mostly for the rent of the theatre (a court for jeu de paume), for which they owed 2000 livres. Historians differ as to whether his father or the lover of a member of his troupe paid his debts; either way, after a 24-hour stint in prison he returned to the acting circuit. It was at this time that he began to use the pseudonym Molière, possibly inspired by a small village of the same name in the Midi near Le Vigan. It was likely that he changed his name to spare his father the shame of having an actor in the family (actors, although no longer vilified by the state under Louis XIV, were still not allowed to be buried in sacred ground).

After his imprisonment, he and Madeleine began a theatrical circuit of the provinces with a new theatre troupe; this life was to last about twelve years, during which he initially played in the company of Charles Dufresne, and subsequently created a company of his own, which had sufficient success and obtained the patronage of Philippe I, Duke of Orléans. Few plays survive from this period. The most noteworthy are L'Étourdi ou les Contretemps (The Bungler) and Le Docteur Amoureux (The Doctor in Love); with these two plays, Molière moved away from the heavy influence of the Italian improvisational Commedia dell'arte, and displayed his talent for mockery. In the course of his travels he met Armand, Prince of Conti, the governor of Languedoc, who became his patron, and named his company after him. This friendship later ended when Armand, having contracted syphilis from a courtesan, turned toward religion and joined Molière's enemies in the Parti des Dévots and the Compagnie de Saint Sacrement.

In Lyon, Mademoiselle Du Parc, known as Marquise, joined the company. Marquise was courted, in vain, by Pierre Corneille and later became the lover of Jean Racine. Racine offered Molière his tragedy Théagène et Chariclée (one of the early works he wrote after he had abandoned his theology studies), but Molière would not perform it, though he encouraged Racine to pursue his artistic career.

Return to Paris
Molière was forced to reach Paris in stages, staying outside for a few weeks in order to promote himself with society gentlemen and allow his reputation to feed in to Paris. Molière reached Paris in 1658 and performed in front of the King at the Louvre (then for rent as a theatre) in Corneille's tragedy Nicomède and in the farce Le Docteur Amoureux with some success. He was awarded the title of Troupe de Monsieur (Monsieur being the honorific for the king's brother Philippe I, Duke of Orléans). With the help of Monsieur, his company was allowed to share the theatre in the large hall of the Petit-Bourbon with the Italian Commedia dell'arte company of Tiberio Fiorillo, famous for the character of Scaramouche. (The two companies performed in the theatre on different nights.) The premiere of Molière's Les Précieuses Ridicules (The Affected Young Ladies) took place at the Petit-Bourbon on 18 November 1659.

Les Précieuses Ridicules was the first of Molière's many attempts to satirize certain societal mannerisms and affectations then common in France. It is widely accepted that the plot was based on Samuel Chappuzeau's Le Cercle des Femmes of 1656. He primarily mocks the Académie Française, a group created by Richelieu under a royal patent to establish the rules of the fledgling French theatre. The Académie preached unity of time, action, and styles of verse. Molière is often associated with the claim that comedy castigat ridendo mores or "criticises customs through humour" (a phrase in fact coined by his contemporary Jean de Santeuil and sometimes mistaken for a classical Latin proverb).

Height of fame
Despite his own preference for tragedy, which he had tried to further with the Illustre Théâtre, Molière became famous for his farces, which were generally in one act and performed after the tragedy. Some of these farces were only partly written, and were played in the style of Commedia dell'arte with improvisation over a canovaccio (a vague plot outline). He began to write full, five-act comedies in verse (L'Étourdi (Lyon, 1654) and Le dépit amoureux (Béziers, 1656)), which although immersed in the gags of contemporary Italian troupes, were successful as part of Madeleine Béjart and Molière's plans to win aristocratic patronage and, ultimately, move the troupe to a position in a Paris theater-venue. Later Molière concentrated on writing musical comedies, in which the drama is interrupted by songs and/or dances, but for years the fundamentals of numerous comedy-traditions would remain strong, especially Italian (e.g. the semi-improvisatory style that in the 1750s writers started calling commedia dell'arte), Spanish, and French plays, all also drawing on classical models (e.g. Plautus and Terence), especially the trope of the clever slave/servant.

Les précieuses ridicules won Molière the attention and the criticism of many, but it was not a popular success. He then asked Fiorillo to teach him the techniques of Commedia dell'arte. His 1660 play Sganarelle, ou Le Cocu imaginaire (The Imaginary Cuckold) seems to be a tribute both to Commedia dell'arte and to his teacher. Its theme of marital relationships dramatizes Molière's pessimistic views on the falsity inherent in human relationships. This view is also evident in his later works and was a source of inspiration for many later authors, including (with different effect), 20th century Nobel Prize winner Luigi Pirandello. It describes a kind of round dance where two couples believe that each of their partners has been betrayed by the other's and is the first in Molière's "Jealousy series", which includes Dom Garcie de Navarre, L'École des maris and L'École des femmes.

In 1660, the Petit-Bourbon was demolished to make way for the eastern expansion of the Louvre, but Molière's company was allowed to move into the abandoned theatre in the east wing of the Palais-Royal. After a period of refurbishment they opened there on 20 January 1661. In order to please his patron, Monsieur, who was so enthralled with entertainment and art that he was soon excluded from state affairs, Molière wrote and played Dom Garcie de Navarre ou Le Prince jaloux (The Jealous Prince, 4 February 1661), a heroic comedy derived from a work of Cicognini. Two other comedies of the same year were the successful L'École des maris (The School for Husbands) and Les Fâcheux (The Bores), subtitled Comédie faite pour les divertissements du Roi (a comedy for the King's amusements) because it was performed during a series of parties that Nicolas Fouquet gave in honor of the sovereign. These entertainments led Jean-Baptiste Colbert to demand the arrest of Fouquet for wasting public money, and he was condemned to life imprisonment.

On 20 February 1662, Molière married Armande Béjart, whom he believed to be the sister of Madeleine. (She may have been her illegitimate daughter with the Duke of Modena.) The same year, he premiered L'École des femmes (The School for Wives), subsequently regarded as a masterpiece. It poked fun at the limited education that was given to daughters of rich families and reflected Molière's own marriage. Both this work and his marriage attracted much criticism. The play sparked the protest called the "Quarrel of L'École des femmes". On the artistic side he responded with two lesser-known works: La Critique de "L'École des femmes", in which he imagined the spectators of his previous work attending it. The piece mocks the people who had criticised L'École des femmes by showing them at dinner after watching the play; it addresses all the criticism raised about the piece by presenting the critics' arguments and then dismissing them. This was the so-called Guerre comique (War of Comedy), in which the opposite side was taken by writers like Donneau de Visé, Edmé Boursault, and Montfleury.

However, more serious opposition was brewing, focusing on Molière's politics and his personal life. A so-called parti des Dévots arose in French high society, who protested against Molière's excessive "realism" and irreverence, which were causing some embarrassment. These people accused Molière of having married his daughter. The Prince of Conti, once Molière's friend, joined them. Molière had other enemies, too, among them the Jansenists and some traditional authors. However, the king expressed support for the Moliere, granting him a pension and agreeing to be the godfather of Molière's first son. Boileau also supported him through statements that he included in his Art poétique.

Molière's friendship with Jean-Baptiste Lully influenced him towards writing his Le Mariage forcé and La Princesse d'Élide (subtitled as Comédie galante mêlée de musique et d'entrées de ballet), written for royal "divertissements" at the Palace of Versailles.

Tartuffe, ou L'Imposteur was also performed at Versailles, in 1664, and created the greatest scandal of Molière's artistic career. Its depiction of the hypocrisy of the dominant classes was taken as an outrage and violently contested. It also aroused the wrath of the Jansenists and the play was banned.

Molière was always careful not to attack the institution of monarchy. He earned a position as one of the king's favourites and enjoyed his protection from the attacks of the court. The king allegedly suggested that Molière suspend performances of Tartuffe, and the author rapidly wrote Dom Juan ou le Festin de Pierre to replace it. It was a strange work, derived from a work by Tirso de Molina and rendered in a prose that still seems modern today. It describes the story of an atheist who becomes a religious hypocrite and, for this, is punished by God. This work too was quickly suspended. The king, demonstrating his protection once again, became the new official sponsor of Molière's troupe.

With music by Lully, Molière presented L'Amour médecin (Love Doctor or Medical Love). Subtitles on this occasion reported that the work was given "par ordre du Roi" (by order of the king) and this work was received much more warmly than its predecessors.

In 1666, Le Misanthrope was produced. It is now widely regarded as Molière's most refined masterpiece, the one with the highest moral content, but it was little appreciated at the time. It caused the "conversion" of Donneau de Visé, who became fond of his theatre. But it was a commercial flop, forcing Molière to immediately write Le médecin malgré lui (The Doctor Despite Himself), a satire against the official sciences. This was a success despite a moral treatise by the Prince of Conti, criticizing the theatre in general and Molière in particular. In several of his plays, Molière depicted the physicians of his day as pompous individuals who speak (poor) Latin to impress others with false erudition, and know only clysters and bleedings as (ineffective) remedies.

After the Mélicerte and the Pastorale comique, he tried again to perform a revised Tartuffe in 1667, this time with the name of Panulphe or L'Imposteur. As soon as the King left Paris for a tour, Lamoignon and the archbishop banned the play. The King finally imposed respect for Tartuffe a few years later, after he had gained more power over the clergy.

Molière, now ill, wrote less. Le Sicilien ou L'Amour peintre was written for festivities at the castle of Saint-Germain-en-Laye, and was followed in 1668 by Amphitryon, inspired both by Plautus' work of the same name and Jean Rotrou's successful reconfiguration of the drama. With some conjecture, Molière's play can be seen to allude to the love affairs of Louis XIV, then king of France. George Dandin, ou Le mari confondu (The Confounded Husband) was little appreciated, but success returned with L'Avare (The Miser), now very well known.

With Lully, he again used music for Monsieur de Pourceaugnac, for Les Amants magnifiques, and finally for Le Bourgeois gentilhomme (The Middle Class Gentleman), another of his masterpieces. It is claimed to be particularly directed against Colbert, the minister who had condemned his old patron Fouquet. The collaboration with Lully ended with a tragédie et ballet, Psyché, written in collaboration with Pierre Corneille and Philippe Quinault.

In 1672, Madeleine Béjart died, and Molière suffered from this loss and from the worsening of his own illness. Nevertheless, he wrote a successful Les Fourberies de Scapin ("Scapin's Deceits"), a farce and a comedy in five acts. His following play, La Comtesse d'Escarbagnas, is considered one of his lesser works.

Les Femmes savantes (The Learned Ladies) of 1672 is considered another of Molière's masterpieces. It was born from the termination of the legal use of music in theatre, since Lully had patented the opera in France (and taken most of the best available singers for his own performances), so Molière had to go back to his traditional genre. It was a great success, and it led to his last work, which is still held in high esteem.

In his 14 years in Paris, Molière single-handedly wrote 31 of the 85 plays performed on his stage.

Les Comédies-Ballets
In 1661, Molière introduced the comédies-ballets in conjunction with Les Fâcheux. These ballets were a transitional form of dance performance between the court ballets of Louis XIV and the art of professional theatre which was developing in the advent of the use of the proscenium stage. The comédies-ballets developed accidentally when Molière was enlisted to mount both a play and a ballet in the honor of Louis XIV and found that he did not have a big enough cast to meet these demands. Molière therefore decided to combine the ballet and the play so that his goal could be met while the performers catch their breath and change costume. The risky move paid off and Molière was asked to produce twelve more comédies-ballets before his death. During the comédies-ballets, Molière collaborated with Pierre Beauchamp. who codified the five balletic positions of the feet and arms and was partly responsible for the creation of the Beauchamp-Feuillet dance notation. Molière also collaborated with Jean-Baptiste Lully. Lully was a dancer, choreographer, and composer, whose dominant reign at the Paris Opéra lasted 15 years. Under his command, ballet and opera rightly became professional arts unto themselves. The comédies-ballets closely integrated dance with music and the action of the play and the style of continuity distinctly separated these performances from the court ballets of the time; additionally, the comédies-ballets demanded that both the dancers and the actors play an important role in advancing the story. Similar to the court ballets, both professionally trained dancers and courtiers socialized together at the comédies-ballets - Louis XIV even played the part of an Egyptian in Molière's Le Mariage forcé (1664) and also appeared as Neptune and Apollo in his retirement performance of Les Amants magnifiques (1670).

Death
Molière suffered from pulmonary tuberculosis, possibly contracted when he was imprisoned for debt as a young man. The circumstances of Molière's death, on 17 February 1673, became legend. He collapsed on stage in a fit of coughing and haemorrhaging while performing in the last play he had written, which had lavish ballets performed to the music of Marc-Antoine Charpentier and which ironically was titled Le Malade imaginaire (The Imaginary Invalid). Molière insisted on completing his performance. Afterwards he collapsed again with another, larger haemorrhage before being taken home, where he died a few hours later, without receiving the last rites because two priests refused to visit him while a third arrived too late. The superstition that green brings bad luck to actors is said to originate from the colour of the clothing he was wearing at the time of his death.

Under French law at the time, actors were not allowed to be buried in the sacred ground of a cemetery. However, Molière's widow, Armande, asked the King if her spouse could be granted a normal funeral at night. The King agreed and Molière's body was buried in the part of the cemetery reserved for unbaptised infants.

In 1792, his remains were brought to the museum of French monuments, and in 1817, transferred to Père Lachaise Cemetery in Paris, close to those of La Fontaine.

Reception of his works

Though conventional thinkers, religious leaders and medical professionals in Molière's time criticised his work, their ideas did not really diminish his widespread success with the public. Other playwrights and companies began to emulate his dramatic style in England and in France. Molière's works continued to garner positive feedback in 18th-century England, but they were not so warmly welcomed in France at this time. However, during the French Restoration of the 19th century, Molière's comedies became popular with both the French public and the critics. Romanticists admired his plays for the unconventional individualism they portrayed. 20th-century scholars have carried on this interest in Molière and his plays and have continued to study a wide array of issues relating to this playwright. Many critics now are shifting their attention from the philosophical, religious and moral implications in his comedies to the study of his comic technique.

Molière's works were translated into English prose by John Ozell in 1714, but the first complete version in English, by Baker and Miller in 1739, remained "influential" and was long reprinted. The first to offer full translations of Molière's verse plays such as Tartuffe into English verse was Curtis Hidden Page, who produced blank verse versions of three of the plays in his 1908 translation. Since then, notable translations have been made by Richard Wilbur, Donald M. Frame, and many others.

In his memoir A Terrible Liar, actor Hume Cronyn writes that, in 1962, celebrated actor Laurence Olivier criticized Molière. According to Cronyn, he mentioned to Olivier that he (Cronyn) was about to play the title role in The Miser, and that Olivier then responded "Molière? Funny as a baby's open grave." Cronyn comments on the incident: "You may imagine how that made me feel. Fortunately, he was dead wrong."

Author Martha Bellinger points out that:

"[Molière] has been accused of not having a consistent, organic style, of using faulty grammar, of mixing his metaphors, and of using unnecessary words for the purpose of filling out his lines. All these things are occasionally true, but they are trifles in comparison to the wealth of character he portrayed, to his brilliancy of wit, and to the resourcefulness of his technique. He was wary of sensibility or pathos; but in place of pathos he had "melancholy — a puissant and searching melancholy, which strangely sustains his inexhaustible mirth and his triumphant gaiety"."

Influence on French culture
Molière is considered the creator of modern French comedy. Many words or phrases introduced in Molière's plays are still used in current French:
 * A tartuffe is a hypocrite, especially a hypocrite displaying affected morality or religious piety.
 * A harpagon, named after the main character of The Miser, is an obsessively greedy and cheap man.
 * The statue of the Commander (statue du Commandeur) from Dom Juan is used as a model of implacable rigidity (raide comme la statue du Commandeur).
 * In Les Fourberies de Scapin, Act II, scene 7, Géronte is asked for ransom money for his son, allegedly held in a galley. He repeats, "What the deuce did he want to go into that galley for?" (Que diable allait-il faire dans cette galère?) The phrase "to go into that galley" is used to describe unnecessary difficulties a person has sought, and galère ("galley") means a difficult and chaotic situation.
 * In Tartuffe, act 3, scene 2, Tartuffe insists that Dorine take a handkerchief to cover up her bosom, saying, "Cover that bosom which I ought not to see" (Couvrez ce sein que je ne saurais voir). This phrase (often with cachez, "hide," instead of couvrez, and often with some other item replacing sein) is frequently used to imply that someone else is calling for something to be hidden or ignored out of their own hypocrisy, disingenuousness, censoriousness, etc.
 * In Le médecin malgré lui, forced to impersonate a doctor, the chancer Sganarelle examines a young woman who is faking muteness in order to delay an arranged marriage. He then delivers to her father a "diagnosis" which consists of strings of gibberish, dog latin and recursive explanations which conclude with an authoritative "and so that is why your daughter is mute" (Et voilà pourquoi votre fille est muette). The phrase is used wholesale to mock an unsatisfactory explanation.
 * Monsieur Jourdain in Le Bourgeois gentilhomme arranges to be tutored in good manners and culture, and is delighted to learn that, because every statement that is not poetry is prose, he therefore has been speaking prose for 40 years without knowing it (Par ma foi, il y a plus de quarante ans que je dis de la prose, sans que j’en susse rien). The more modern phrase "je parle de la prose sans le savoir" is used by a person who realizes that he was more skilled or better aligned than he thought.
 * In the Comédie-ballet "George Dandin" (1668), Act I, scene 7, the main character uses the phrase Tu l'as voulu, George Dandin ("You wanted it, George Dandin") to address himself when his rich wife cheats on him. Now the phrase is used to reproach someone ironically, something like "You did it yourself".

Portrayals of Molière
Molière plays a small part in Alexandre Dumas's novel The Vicomte of Bragelonne, in which he is seen taking inspiration from the musketeer Porthos for his central character in Le Bourgeois gentilhomme.

Russian writer Mikhail Bulgakov wrote a semi-fictitious biography-tribute to Molière, titled Life of Mr. de Molière. It was written in 1932–1933 and first published 1962.

The French 1978 film simply titled Molière directed by Ariane Mnouchkine and starring Philippe Caubère presents his complete biography. It was in competition for the Palme d'Or at Cannes in 1978.

He is portrayed among other writers in The Blasphemers' Banquet (1989).

The 2000 film Le Roi Danse (The King Dances), in which Molière is played by Tchéky Karyo, shows his collaborations with Jean-Baptiste Lully, as well as his illness and on-stage death.

The 2007 French film Molière was more loosely based on the life of Molière, starring Romain Duris, Fabrice Luchini and Ludivine Sagnier.

David Hirson's play La Bête, written in the style of Molière, includes the character Elomire as an anagrammatic parody of him.

The 2023 musical Molière, l'Opéra Urbain, directed by Bruno Berberes and staged at the Dôme de Paris from November 11, 2023 to February 18, 2024, is a retelling of the life of Molière using a blend of historical costuming with contemporary artistic styles in staging and musical genres.

List of major works

 * Le Médecin volant (1645)—The Flying Doctor
 * La Jalousie du barbouillé (1650)—The Jealousy of le Barbouillé
 * L'Étourdi ou les Contretemps (1655)—The Blunderer, or, the Counterplots
 * Le Dépit amoureux (16 December 1656)—The Love-Tiff
 * Le Docteur amoureux (1658), the first play performed by Molière's troupe for Louis XIV (now lost)—The Doctor in Love
 * Les Précieuses ridicules (18 November 1659)—The Affected Young Ladies
 * Sganarelle ou Le Cocu imaginaire (28 May 1660)—Sganarelle, or the Imaginary Cuckold
 * Dom Garcie de Navarre ou Le Prince jaloux (4 February 1661)—Don Garcia of Navarre or the Jealous Prince
 * L'École des maris (24 June 1661)—The School for Husbands
 * Les Fâcheux (17 August 1661)—The Bores (also translated The Mad)
 * L'École des femmes (26 December 1662; adapted into The Amorous Flea, 1964)—The School for Wives
 * La Jalousie du Gros-René (15 April 1663; now lost)—The Jealousy of Gros-René
 * La Critique de l'école des femmes (1 June 1663)—Critique of the School for Wives
 * L'Impromptu de Versailles (14 October 1663)—The Versailles Impromptu
 * Le Mariage forcé (29 January 1664)—The Forced Marriage
 * Gros-René, petit enfant  (27 April 1664; now lost)—Gros-René, Small Child
 * La Princesse d'Élide (8 May 1664)—The Princess of Elid
 * Tartuffe ou L'Imposteur (12 May 1664)—Tartuffe, or, the Impostor
 * Dom Juan ou Le Festin de pierre (15 February 1665)—Don Juan, or, The Stone Banquet (subtitle also translated The Stone Guest, The Feast with the Statue, &c.)
 * L'Amour médecin (15 September 1665)—Love Is the Doctor
 * Le Misanthrope ou L'Atrabilaire amoureux (4 June 1666)—The Misanthrope, or, the Cantankerous Lover
 * Le Médecin malgré lui (6 August 1666)—The Doctor in Spite of Himself
 * Mélicerte (2 December 1666)
 * Pastorale comique (5 January 1667)—Comic Pastoral
 * Le Sicilien ou L'Amour peintre (14 February 1667)—The Sicilian, or Love the Painter
 * Amphitryon (13 January 1668)
 * George Dandin ou Le Mari confondu (18 July 1668)—George Dandin, or the Abashed Husband
 * L'Avare ou L'École du mensonge (9 September 1668)—The Miser, or, the School for Lies
 * Monsieur de Pourceaugnac (6 October 1669)
 * Les Amants magnifiques (4 February 1670)—The Magnificent Lovers
 * Le Bourgeois gentilhomme (14 October 1670)—The Bourgeois Gentleman
 * Psyché (17 January 1671)—Psyche
 * Les Fourberies de Scapin (24 May 1671)—The Impostures of Scapin
 * La Comtesse d'Escarbagnas (2 December 1671)—The Countess of Escarbagnas
 * Les Femmes savantes (11 March 1672)—The Learned Ladies
 * Le Malade imaginaire (10 February 1673)—The Imaginary Invalid (or The Hypochondriac)

title: Molière


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Mold

A mold (US, PH) or mould (UK, CW) is one of the structures that certain fungi can form. The dust-like, colored appearance of molds is due to the formation of spores containing fungal secondary metabolites. The spores are the dispersal units of the fungi. Not all fungi form molds. Some fungi form mushrooms; others grow as single cells and are called microfungi (for example yeasts).

A large and taxonomically diverse number of fungal species form molds. The growth of hyphae results in discoloration and a fuzzy appearance, especially on food. The network of these tubular branching hyphae, called a mycelium, is considered a single organism. The hyphae are generally transparent, so the mycelium appears like very fine, fluffy white threads over the surface. Cross-walls (septa) may delimit connected compartments along the hyphae, each containing one or multiple, genetically identical nuclei. The dusty texture of many molds is caused by profuse production of asexual spores (conidia) formed by differentiation at the ends of hyphae. The mode of formation and shape of these spores is traditionally used to classify molds. Many of these spores are colored, making the fungus much more obvious to the human eye at this stage in its life-cycle.

Molds are considered to be microbes and do not form a specific taxonomic or phylogenetic grouping, but can be found in the divisions Zygomycota and Ascomycota. In the past, most molds were classified within the Deuteromycota. Mold had been used as a common name for now non-fungal groups such as water molds or slime molds that were once considered fungi.

Molds cause biodegradation of natural materials, which can be unwanted when it becomes food spoilage or damage to property. They also play important roles in biotechnology and food science in the production of various pigments, foods, beverages, antibiotics, pharmaceuticals and enzymes. Some diseases of animals and humans can be caused by certain molds: disease may result from allergic sensitivity to mold spores, from growth of pathogenic molds within the body, or from the effects of ingested or inhaled toxic compounds (mycotoxins) produced by molds.

Biology
There are thousands of known species of mold fungi with diverse life-styles including saprotrophs, mesophiles, psychrophiles and thermophiles, and a very few opportunistic pathogens of humans. They all require moisture for growth and some live in aquatic environments. Like all fungi, molds derive energy not through photosynthesis but from the organic matter on which they live, utilizing heterotrophy. Typically, molds secrete hydrolytic enzymes, mainly from the hyphal tips. These enzymes degrade complex biopolymers such as starch, cellulose and lignin into simpler substances which can be absorbed by the hyphae. In this way, molds play a major role in causing decomposition of organic material, enabling the recycling of nutrients throughout ecosystems. Many molds also synthesize mycotoxins and siderophores which, together with lytic enzymes, inhibit the growth of competing microorganisms. Molds can also grow on stored food for animals and humans, making the food unpalatable or toxic and are thus a major source of food losses and illness. Many strategies for food preservation (salting, pickling, jams, bottling, freezing, drying) are to prevent or slow mold growth as well as the growth of other microbes.

Molds reproduce by producing large numbers of small spores, which may contain a single nucleus or be multinucleate. Mold spores can be asexual (the products of mitosis) or sexual (the products of meiosis); many species can produce both types. Some molds produce small, hydrophobic spores that are adapted for wind dispersal and may remain airborne for long periods; in some the cell walls are darkly pigmented, providing resistance to damage by ultraviolet radiation. Other mold spores have slimy sheaths and are more suited to water dispersal. Mold spores are often spherical or ovoid single cells, but can be multicellular and variously shaped. Spores may cling to clothing or fur; some are able to survive extremes of temperature and pressure.

Although molds can grow on dead organic matter everywhere in nature, their presence is visible to the unaided eye only when they form large colonies. A mold colony does not consist of discrete organisms but is an interconnected network of hyphae called a mycelium. All growth occurs at hyphal tips, with cytoplasm and organelles flowing forwards as the hyphae advance over or through new food sources. Nutrients are absorbed at the hyphal tip. In artificial environments such as buildings, humidity and temperature are often stable enough to foster the growth of mold colonies, commonly seen as a downy or furry coating growing on food or other surfaces.

Few molds can begin growing at temperatures of 4 C or below, so food is typically refrigerated at this temperature. When conditions do not enable growth to take place, molds may remain alive in a dormant state depending on the species, within a large range of temperatures. The many different mold species vary enormously in their tolerance to temperature and humidity extremes. Certain molds can survive harsh conditions such as the snow-covered soils of Antarctica, refrigeration, highly acidic solvents, anti-bacterial soap and even petroleum products such as jet fuel.

Xerophilic molds are able to grow in relatively dry, salty, or sugary environments, where water activity (aw) is less than 0.85; other molds need more moisture.

Common molds
Common genera of molds include:

 * Acremonium
 * Alternaria
 * Aspergillus
 * Cladosporium
 * Fusarium
 * Mucor
 * Penicillium
 * Rhizopus
 * Stachybotrys
 * Trichoderma
 * Trichophyton

Food production
The Kōji molds are a group of Aspergillus species, notably Aspergillus oryzae, and secondarily A. sojae, that have been cultured in eastern Asia for many centuries. They are used to ferment a soybean and wheat mixture to make soybean paste and soy sauce. Koji molds break down the starch in rice, barley, sweet potatoes, etc., a process called saccharification, in the production of sake, shōchū and other distilled spirits. Koji molds are also used in the preparation of Katsuobushi.

Red rice yeast is a product of the mold Monascus purpureus grown on rice, and is common in Asian diets. The yeast contains several compounds collectively known as monacolins, which are known to inhibit cholesterol synthesis. A study has shown that red rice yeast used as a dietary supplement, combined with fish oil and healthy lifestyle changes, may help reduce "bad" cholesterol as effectively as certain commercial statin drugs. Nonetheless, other work has shown it may not be reliable (perhaps due to non-standardization) and even toxic to liver and kidneys.

Some sausages, such as salami, incorporate starter cultures of molds to improve flavor and reduce bacterial spoilage during curing. Penicillium nalgiovense, for example, may appear as a powdery white coating on some varieties of dry-cured sausage.

Other molds that have been used in food production include:
 * Fusarium venenatum – quorn
 * Geotrichum candidum – cheese
 * Neurospora sitophila – oncom
 * Penicillium spp. – various cheeses including Brie and Blue cheese
 * Rhizomucor miehei – microbial rennet for making vegetarian and other cheeses
 * Rhizopus oligosporus – tempeh
 * Rhizopus oryzae – tempeh, jiuqu for jiuniang or precursor for making Chinese rice wine

Pharmaceuticals from molds
Alexander Fleming's accidental discovery of the antibiotic penicillin involved a Penicillium mold called Penicillium rubrum (although the species was later established to be Penicillium rubens). Fleming continued to investigate penicillin, showing that it could inhibit various types of bacteria found in infections and other ailments, but he was unable to produce the compound in large enough amounts necessary for production of a medicine. His work was expanded by a team at Oxford University; Clutterbuck, Lovell, and Raistrick, who began to work on the problem in 1931. This team was also unable to produce the pure compound in any large amount, and found that the purification process diminished its effectiveness and negated the anti-bacterial properties it had.

Howard Florey, Ernst Chain, Norman Heatley, Edward Abraham, also all at Oxford, continued the work. They enhanced and developed the concentration technique by using organic solutions rather than water, and created the "Oxford Unit" to measure penicillin concentration within a solution. They managed to purify the solution, increasing its concentration by 45–50 times, but found that a higher concentration was possible. Experiments were conducted and the results published in 1941, though the quantities of penicillin produced were not always high enough for the treatments required. As this was during the Second World War, Florey sought US government involvement. With research teams in the UK and some in the US, industrial-scale production of crystallized penicillin was developed during 1941–1944 by the USDA and by Pfizer.

Several statin cholesterol-lowering drugs (such as lovastatin, from Aspergillus terreus) are derived from molds.

The immunosuppressant drug cyclosporine, used to suppress the rejection of transplanted organs, is derived from the mold Tolypocladium inflatum.

Health effects
Molds are ubiquitous, and mold spores are a common component of household and workplace dust; however, when mold spores are present in large quantities, they can present a health hazard to humans, potentially causing allergic reactions and respiratory problems.

Some molds also produce mycotoxins that can pose serious health risks to humans and animals. Some studies claim that exposure to high levels of mycotoxins can lead to neurological problems and, in some cases, death. Prolonged exposure, e.g. daily home exposure, may be particularly harmful. Research on the health impacts of mold has not been conclusive. The term "toxic mold" refers to molds that produce mycotoxins, such as Stachybotrys chartarum, and not to all molds in general. Mold in the home can usually be found in damp, dark or steamy areas, e.g. bathrooms, kitchens, cluttered storage areas, recently flooded areas, basement areas, plumbing spaces, areas with poor ventilation and outdoors in humid environments. Symptoms caused by mold allergy are: watery, itchy eyes; a chronic cough; headaches or migraines; difficulty breathing; rashes; tiredness; sinus problems; nasal blockage and frequent sneezing.

Molds can also pose a hazard to human and animal health when they are consumed following the growth of certain mold species in stored food. Some species produce toxic secondary metabolites, collectively termed mycotoxins, including aflatoxins, ochratoxins, fumonisins, trichothecenes, citrinin, and patulin. These toxic properties may be used for the benefit of humans when the toxicity is directed against other organisms; for example, penicillin adversely affects the growth of Gram-positive bacteria (e.g. Clostridium species), certain spirochetes and certain fungi.

Growth in buildings and homes
Mold growth in buildings generally occurs as fungi colonize porous building materials, such as wood. Many building products commonly incorporate paper, wood products, or solid wood members, such as paper-covered drywall, wood cabinets, and insulation. Interior mold colonization can lead to a variety of health problems as microscopic airborne reproductive spores, analogous to tree pollen, are inhaled by building occupants. High quantities of indoor airborne spores as compared to exterior conditions are strongly suggestive of indoor mold growth. Determination of airborne spore counts is accomplished by way of an air sample, in which a specialized pump with a known flow rate is operated for a known period of time. To account for background levels, air samples should be drawn from the affected area, a control area, and the exterior.

The air sampler pump draws in air and deposits microscopic airborne particles on a culture medium. The medium is cultured in a laboratory and the fungal genus and species are determined by visual microscopic observation. Laboratory results also quantify fungal growth by way of a spore count for comparison among samples. The pump operation time is recorded and when multiplied by pump flow rate results in a specific volume of air obtained. Although a small volume of air is actually analyzed, common laboratory reports extrapolate the spore count data to estimate spores that would be present in a cubic meter of air.

Mold spores are drawn to specific environments, making it easier for them to grow. These spores will usually only turn into a full-blown outbreak if certain conditions are met. Various practices can be followed to mitigate mold issues in buildings, the most important of which is to reduce moisture levels that can facilitate mold growth. Air filtration reduces the number of spores available for germination, especially when a High Efficiency Particulate Air (HEPA) filter is used. A properly functioning AC unit also reduces the relative humidity in rooms. The United States Environmental Protection Agency (EPA) currently recommends that relative humidity be maintained below 60%, ideally between 30% and 50%, to inhibit mold growth.

Eliminating the moisture source is the first step at fungal remediation. Removal of affected materials may also be necessary for remediation, if materials are easily replaceable and not part of the load-bearing structure. Professional drying of concealed wall cavities and enclosed spaces such as cabinet toekick spaces may be required. Post-remediation verification of moisture content and fungal growth is required for successful remediation. Many contractors perform post-remediation verification themselves, but property owners may benefit from independent verification.

Use in art
Various artists have used mold in various artistic fashions. Daniele Del Nero, for example, constructs scale models of houses and office buildings and then induces mold to grow on them, giving them an unsettling, reclaimed-by-nature look. Stacy Levy sandblasts enlarged images of mold onto glass, then allows mold to grow in the crevasses she has made, creating a macro-micro portrait. Sam Taylor-Johnson, director of Nowhere Boy and Fifty Shades of Grey (among others), has made a number of time-lapse films capturing the gradual decay of classically arranged still lifes.

license: Creative Commons - Attribution Share-Alike - https://creativecommons.org/licenses/by-sa/4.0/

title: Mold


text:
Mycelium

Mycelium (: mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Its normal form is that of branched, slender, entangled, anastomosing, hyaline threads. Fungal colonies composed of mycelium are found in and on soil and many other substrates. A typical single spore germinates into a monokaryotic mycelium, which cannot reproduce sexually; when two compatible monokaryotic mycelia join and form a dikaryotic mycelium, that mycelium may form fruiting bodies such as mushrooms. A mycelium may be minute, forming a colony that is too small to see, or may grow to span thousands of acres as in Armillaria.

Through the mycelium, a fungus absorbs nutrients from its environment. It does this in a two-stage process. First, the hyphae secrete enzymes onto or into the food source, which break down biological polymers into smaller units such as monomers. These monomers are then absorbed into the mycelium by facilitated diffusion and active transport.

Mycelia are vital in terrestrial and aquatic ecosystems for their role in the decomposition of plant material. They contribute to the organic fraction of soil, and their growth releases carbon dioxide back into the atmosphere (see carbon cycle). Ectomycorrhizal extramatrical mycelium, as well as the mycelium of arbuscular mycorrhizal fungi, increase the efficiency of water and nutrient absorption of most plants and confers resistance to some plant pathogens. Mycelium is an important food source for many soil invertebrates. They are vital to agriculture and are important to almost all species of plants, many species co-evolving with the fungi. Mycelium is a primary factor in some plants’ health, nutrient intake and growth, with mycelium being a major factor to plant fitness.

Networks of mycelia can transport water and spikes of electrical potential.

Sclerotia are compact or hard masses of mycelium.

Agriculture
One of the primary roles of fungi in an ecosystem is to decompose organic compounds. Petroleum products and some pesticides (typical soil contaminants) are organic molecules (i.e., they are built on a carbon structure), and thereby show a potential carbon source for fungi. Hence, fungi have the potential to eradicate such pollutants from their environment unless the chemicals prove toxic to the fungus. This biological degradation is a process known as bioremediation.

Mycelial mats have been suggested as having potential as biological filters, removing chemicals and microorganisms from soil and water. The use of fungal mycelium to accomplish this has been termed mycofiltration.

Knowledge of the relationship between mycorrhizal fungi and plants suggests new ways to improve crop yields.

When spread on logging roads, mycelium can act as a binder, holding disturbed new soil in place preventing washouts until woody plants can establish roots.

Fungi are essential for converting biomass into compost, as they decompose feedstock components such as lignin, which many other composting microorganisms cannot. Turning a backyard compost pile will commonly expose visible networks of mycelia that have formed on the decaying organic material within. Compost is an essential soil amendment and fertilizer for organic farming and gardening. Composting can divert a substantial fraction of municipal solid waste from landfills.

Commercial
Alternatives to polystyrene and plastic packaging can be produced by growing mycelium in agricultural waste.

Mycelium has also been used as a material in furniture, and artificial leather.

One of the main commercial uses of mycelium is its use to create artificial leather. Animal leather contributes to a significant environmental footprint, as livestock farming is associated with deforestation, greenhouse gas emissions, and grazing. In addition, the production of synthetic leathers from polyvinyl chloride and polyurethane require the use of hazardous chemicals and fossil fuels, and they are not biodegradable (like plastic). Fungal-based artificial leather is cheaper to produce, has less of an environmental footprint, and is biodegradable. It costs between 18 and 28 cents to produce a square meter of raw mycelium, while it costs between $5.81 and $6.24 to produce a square meter of raw animal hide. Fungal growth is carbon neutral and pure mycelium is 94% biodegradable. However, the use of polymeric materials such as polyester or polylactic acid to improve artificial leather’s properties can negatively affect the biodegradability of the material.

To create leather, fungal mycelium is grown either using liquid-state or solid-state fermentation. In liquid-state fermentation, companies typically use laboratory media or agricultural byproducts to grow fungal biomass. The fungal biomass is then separated into fibers and processed using fiber suspension, filtration, pressing, and drying. These techniques are also commonly utilized in traditional papermaking processes. In solid-state fermentation, mycelium is grown on forestry bioproducts, like sawdust, in an environment with high carbon dioxide concentrations and controlled humidity and temperature. The mycelium mat formed on top of the particle bed is dehydrated, chemically treated, and then compressed to a desired thickness and engraved with a pattern.

Construction material
Mycelium is a strong candidate for sustainable construction primarily due to its lightweight biodegradable structure and its capacity to be grown from waste sources. In addition to this, mycelium has a relatively high strength-to-weight ratio and a much lower embodied energy compared to traditional building materials. Because mycelium takes the form of any mold it's grown in, it can also be advantageous for customization purposes, especially if it's employed as an architectural or aesthetic feature. Current research has also indicated that mycelium does not release toxic resins in the event of a fire because it has a charring effect similar to mass timber. Mycelium plays an interesting role in acoustic insulation, boasting of an absorbance of 70–75% for frequencies of 1500 Hz or less.

Strengths and weaknesses
Mycelium bio-composites have shown strong potential for structural applications, with much higher strength-to-weight ratios than that of conventional materials due primarily to its low density. Compared to conventional building materials, mycelium also has a number of desirable properties that make it an attractive alternative. For example, it has low thermal conductivity and can provide high acoustic insulation. It is biodegradable, has much lower embodied energy, and can serve as a carbon sink, which makes mycelium bio-composites a possible solution to the emissions, energy, and waste associated with building construction.

While mycelium proposes interesting implications as a structural material, there are several significant disadvantages that make it difficult to be practically implemented in large-scale projects. For one, mycelium does not have particularly high compressive strength on its own, ranging from 0.1-0.2 MPa. This is in stark comparison to traditional concrete, which typically has a compressive strength of 17-28 MPa. Even more, because mycelium is considered a living material, it holds specific requirements that make it susceptible to environmental conditions. For instance, it requires a constant source of air in order to stay alive, needs a relatively humid habitat to grow, and cannot be exposed to large amounts of water for fear of contamination and decay.

Mechanical properties
3 separate fungi species (Colorius versicolor, Trametes ochracea, and Ganoderma sessile) were mixed independently with 2 substrates (apple and vine) and tested under separate incubation conditions in order to quantify certain mechanical properties of mycelium. In order to do this, samples were grown in molds, incubated, and dried over the course of 12 days. Samples were tested for water absorption using ASTM C272 guidelines and compared against an EPS material. Tiles of uniform size were cut from the fabricated mold and put under an Instron 3345 machine going at 1 mm/min, up until 20% deformation.

Throughout a 4 stage process, the impact of various substrate and fungal mixes was investigated along with properties of mycelium such as density, water absorption, and compressive strength. Samples were separated into two separate incubation methods and inspected for differences in color, texture, and growth. For the same fungi within each incubation method, minimal differences were recorded. However, across disparate substrate mixtures within the same fungi, colorization and external growth varied between the test samples. While loss of organic matter was calculated, no uniform correlation was found between substrate used and chemical properties of the material. For each of the substrate-fungi mixtures, average densities ranged from 174.1 kg/m3 to 244.9 kg/m3, with the Ganoderma sessile fungi and apple substrate combination being the most dense. Compression tests revealed the Ganoderma sessile fungi and vine substrate to have the highest strength of the samples tested, but no numerical value was provided. For reference, surrounding literature has provided a ballpark estimate of 1-72 kPa. Beyond this, mycelium has a thermal conductivity of 0.05–0.07W/m·K which is less than that of typical concrete.

Construction
The construction of mycelium structures is primarily categorized into three approaches. These include growing blocks in molds, growing in place monolithic structures, and bio-welded units. The first approach cultivates mycelium and its substrate in forms, after which it is dried in ovens and then transported and assembled on site. The second approach uses existing formwork and adapts cast-in-place concrete techniques to grow monolithic mycelium structures in place. The third approach is a hybrid of the previous two referred to as myco-welding, where individual pre-grown units are grown together into a larger monolithic structure.

Studies using grow-in-place methods and myco-welding have explored how to cultivate mycelium and re-use formwork in construction and investigated post-tensioning and friction connections. Research in fabrication has revealed some common challenges faced in construction of mycelium structures, mostly related to the growth of the fungi. It can be difficult to cultivate living material into formwork and it is susceptible to contamination if not properly sterilized. The fungi needs to be kept refrigerated to prevent hardening and properly manage growth and substrate consumption. Additionally, the thickness of fungal growth is limited by the presence of oxygen; if there is no oxygen, the center of the growth can die or be contaminated.

Environmental impact
Researchers have performed life-cycle assessments to evaluate the environmental impact of mycelium bio-composites. Life cycle analysis showed the viability of mycelium as a carbon sink material and as a sustainable alternative to conventional building materials. Use of mycelium as a natural adhesive material may provide environmental benefits, as the fungal-based composites that mycelium is used to create are low cost, low emission, and sustainable. These composites also have a wide range of applications and uses, many of which are in industries responsible for significant environmental pollution, like construction and packaging.

Modern construction and packaging materials are industrially fabricated, non-recyclable, and pollutive: wood products lead to severe deforestation and weather fluctuation; cement is nonbiodegradable and causes high emissions both in production and demolition. Mycelium appears to be cheaper and more sustainable than its counterparts.

Mycelium’s adhesive properties are largely responsible for its diverse array of applications, as it allows them to bind certain substances together. These properties are products of their biological processes, as they secrete corrosive enzymes that allow them to degrade and colonize organic substrates. During degradation, mycelium develops a dense network of thin strands that fuse together within the organic substrate, creating solid material that can hold multiple substrates together. This self-assembly property of mycelium is quite unique, and allows mycelium to grow on a wide range of organic material, including organic waste.

Potential ecological role
Plants appear to communicate within an ecosystem using mycelium, the fungal network produced by mycorrhiza fungi. Mycelial networks constitute 20-30% of soil biomass, though traditional biomass measures fail to detect them. Some 83% of plants appear to exhibit mutualistic association with mycelium as an extension of their root systems, with varying levels of reliance. By some estimates, mycelial networks receive up to 10% or more of the photosynthesis output of their host plants.

This mutualism is initiated by hyphal connections in which mycelial strands infect and attach themselves to plant hyphae, penetrating the cell wall but not entering through the membrane into the plant cytoplasm. Mycelium interacts with the cell at the periarbuscular membrane, which behaves as a sort of exchange medium for nutrients and can produce electrical gradients allowing for electrophysiological signals to be sent and received. In modeling studies, different fungi supply different levels of nutrients and growth-promoting materials, with plants tending to root towards (and thus being infected by) fungi supplying most mineral phosphorus and nitrogen (both essential for plant growth).

Mycorrhizal mycelial associations may intensify competition between individuals of the same species, while alleviating competition between species, via the promotion of inferior competitors, thus promoting plant diversity within its network. In doing so, mycorrhizal fungi promote community ecology, with an added complexity of niche differentiation of different networks and types of mycorrhizal fungi that root at different depths, disperse different organic compounds and nutrients, and have unique interactions with specific species of plants.

Mycelial biology and memory
Several studies have documented the memory capacity of mycelial networks and their adaptability to specific environmental conditions. Mycelia have been specialized for different functions in various climates and develop symbiotic or pathogenic relationships with other organisms, such as the human pathogen Candida auris, which has developed a unique approach of evading detection by human neutrophils through adaptive selection–a process of fungal learning and memory. Additionally, these functions can change based on the scale of the mycelia and nature of the symbiotic relationship; commensal and mutual relationships between fungi and plants form through a separate process known as mycorrhizal association, which are called mycorrhiza. Additionally, hyphal organization into mycelial networks can be deterministic for a variety of functions including biomass retention, water recycling, expansion of future hyphae on a resource efficient approach towards desired nutrient gradients, and the subsequent distribution of these resources across the hyphal network. On a macroscopic scale, many mycelia operate with a sort of hierarchy having a “trunk” or main mycelium, with smaller “branches” branching off. Some saprotrophic basidiomycetes are able to remember past decisions about directional nutrition gradients and will build future mycelium in that direction.

Mycelial memory and intelligence
Current research on collective mycelial intelligence is limited, and while many studies have observed memory and the exchange of electric charge across mycelial networks, this is insufficient evidence to make conclusions about how sensory data is processed in these networks. However, some examples of increased thermal resistance in filamentous fungi suggest a power-law relationship for memory and exposure to a stimulus. Mycelia have also demonstrated the ability to edit their genetic structures within a lifetime due to antibiotic or other extracellular stressors, which can cause rapid acquisition of resistance genes, like those in C. auris. Additionally, plasmodial slime molds demonstrate a similar method of information sharing, as both mycelia and slime molds make use of cAMP molecules for aggregation and signaling.

Sclerotium
Sclerotium is a compact mass of hardened mycelium. For many years, sclerotia were mistaken for individual organisms and described as separate species. However, in the mid 19th century, it was proven that sclerotia was simply a stage in the life cycle of many fungi. Sclerotia are composed of thick, dense shells with dark cells. They are rich in hyphae emergency supplies, such as oil, and they contain small amounts of water. They can survive in dry environments for many years without losing the ability to grow. The size of sclerotia can range from less than a millimeter to tens of centimeters in diameter.

title: Mycelium


text:
Spore

In biology, a spore is a unit of sexual (in fungi) or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. They were thought to have appeared as early as the mid-late Ordovician period as an adaptation of early land plants.

Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula.

In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. In some rare cases, diploid spore is also produced in some algae, or fungi. Under favourable conditions, the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote, which develops into a new sporophyte. This cycle is known as alternation of generations.

The spores of seed plants are produced internally, and the megaspores (formed within the ovules) and the microspores are involved in the formation of more complex structures that form the dispersal units, the seeds and pollen grains.

Definition
The term spore derives from the ancient Greek word σπορά spora, meaning "seed, sowing", related to σπόρος sporos, "sowing", and σπείρειν speirein, "to sow".

In common parlance, the difference between a "spore" and a "gamete" is that a spore will germinate and develop into a sporeling, while a gamete needs to combine with another gamete to form a zygote before developing further.

The main difference between spores and seeds as dispersal units is that spores are unicellular, the first cell of a gametophyte, while seeds contain within them a developing embryo (the multicellular sporophyte of the next generation), produced by the fusion of the male gamete of the pollen tube with the female gamete formed by the megagametophyte within the ovule. Spores germinate to give rise to haploid gametophytes, while seeds germinate to give rise to diploid sporophytes.

Plants
Vascular plant spores are always haploid. Vascular plants are either homosporous (or isosporous) or heterosporous. Plants that are homosporous produce spores of the same size and type.

Heterosporous plants, such as seed plants, spikemosses, quillworts, and ferns of the order Salviniales produce spores of two different sizes: the larger spore (megaspore) in effect functioning as a "female" spore and the smaller (microspore) functioning as a "male". Such plants typically give rise to the two kind of spores from within separate sporangia, either a megasporangium that produces megaspores or a microsporangium that produces microspores. In flowering plants, these sporangia occur within the carpel and anthers, respectively.

Fungi
Fungi commonly produce spores during sexual and asexual reproduction. Spores are usually haploid and grow into mature haploid individuals through mitotic division of cells (Urediniospores and Teliospores among rusts are dikaryotic). Dikaryotic cells result from the fusion of two haploid gamete cells. Among sporogenic dikaryotic cells, karyogamy (the fusion of the two haploid nuclei) occurs to produce a diploid cell. Diploid cells undergo meiosis to produce haploid spores.

Classification of spores
Spores can be classified in several ways such as by their spore producing structure, function, origin during life cycle, and mobility.

Below is a table listing the mode of classification, name, identifying characteristic, examples, and images of different spore species.

External anatomy
Under high magnification, spores often have complex patterns or ornamentation on their exterior surfaces. A specialized terminology has been developed to describe features of such patterns. Some markings represent apertures, places where the tough outer coat of the spore can be penetrated when germination occurs. Spores can be categorized based on the position and number of these markings and apertures. Alete spores show no lines. In monolete spores, there is a single narrow line (laesura) on the spore. Indicating the prior contact of two spores that eventually separated. In trilete spores, each spore shows three narrow lines radiating from a center pole. This shows that four spores shared a common origin and were initially in contact with each other forming a tetrahedron. A wider aperture in the shape of a groove may be termed a colpus. The number of colpi distinguishes major groups of plants. Eudicots have tricolpate spores (i.e. spores with three colpi).

Spore tetrads and trilete spores
Envelope-enclosed spore tetrads are taken as the earliest evidence of plant life on land, dating from the mid-Ordovician (early Llanvirn, ~), a period from which no macrofossils have yet been recovered. Individual trilete spores resembling those of modern cryptogamic plants first appeared in the fossil record at the end of the Ordovician period.

Dispersal

In fungi, both asexual and sexual spores or sporangiospores of many fungal species are actively dispersed by forcible ejection from their reproductive structures. This ejection ensures exit of the spores from the reproductive structures as well as travelling through the air over long distances. Many fungi thereby possess specialized mechanical and physiological mechanisms as well as spore-surface structures, such as hydrophobins, for spore ejection. These mechanisms include, for example, forcible discharge of ascospores enabled by the structure of the ascus and accumulation of osmolytes in the fluids of the ascus that lead to explosive discharge of the ascospores into the air.

The forcible discharge of single spores termed ballistospores involves formation of a small drop of water (Buller's drop), which upon contact with the spore leads to its projectile release with an initial acceleration of more than 10,000 g. Other fungi rely on alternative mechanisms for spore release, such as external mechanical forces, exemplified by puffballs. Attracting insects, such as flies, to fruiting structures, by virtue of their having lively colours and a putrid odour, for dispersal of fungal spores is yet another strategy, most prominently used by the stinkhorns.

In Common Smoothcap moss (Atrichum undulatum), the vibration of sporophyte has been shown to be an important mechanism for spore release.

In the case of spore-shedding vascular plants such as ferns, wind distribution of very light spores provides great capacity for dispersal. Also, spores are less subject to animal predation than seeds because they contain almost no food reserve; however they are more subject to fungal and bacterial predation. Their chief advantage is that, of all forms of progeny, spores require the least energy and materials to produce.

In the spikemoss Selaginella lepidophylla, dispersal is achieved in part by an unusual type of diaspore, a tumbleweed.

Origin
Spores have been found in microfossils dating back to the mid-late Ordovician period. Two hypothesized initial functions of spores relate to whether they appeared before or after land plants. The heavily studied hypothesis is that spores were an adaptation of early land plant species, such as embryophytes, that allowed for plants to easily disperse while adapting to their non-aquatic environment. This is particularly supported by the observation of a thick spore wall in cryptospores. These spore walls would have protected potential offspring from novel weather elements. The second more recent hypothesis is that spores were an early predecessor of land plants and formed during errors in the meiosis of algae, a hypothesized early ancestor of land plants.

Whether spores arose before or after land plants, their contributions to topics in fields like paleontology and plant phylogenetics have been useful. The spores found in microfossils, also known as cryptospores, are well preserved due to the fixed material they are in as well as how abundant and widespread they were during their respective time periods. These microfossils are especially helpful when studying the early periods of earth as macrofossils such as plants are not common nor well preserved. Both cryptospores and modern spores have diverse morphology that indicate possible environmental conditions of earlier periods of Earth and evolutionary relationships of plant species.

title: Spore