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Coccolithophore

Coccolithophores, or coccolithophorids, are single-celled organisms which are part of the phytoplankton, the autotrophic (self-feeding) component of the plankton community. They form a group of about 200 species, and belong either to the kingdom Protista, according to Robert Whittaker's five-kingdom system, or clade Hacrobia, according to a newer biological classification system. Within the Hacrobia, the coccolithophores are in the phylum or division Haptophyta, class Prymnesiophyceae (or Coccolithophyceae). Coccolithophores are almost exclusively marine, are photosynthetic, and exist in large numbers throughout the sunlight zone of the ocean.

Coccolithophores are the most productive calcifying organisms on the planet, covering themselves with a calcium carbonate shell called a coccosphere. However, the reasons they calcify remain elusive. One key function may be that the coccosphere offers protection against microzooplankton predation, which is one of the main causes of phytoplankton death in the ocean.

Coccolithophores are ecologically important, and biogeochemically they play significant roles in the marine biological pump and the carbon cycle. Depending on habitat, they can produce up to 40 percent of the local marine primary production. They are of particular interest to those studying global climate change because, as ocean acidity increases, their coccoliths may become even more important as a carbon sink. Management strategies are being employed to prevent eutrophication-related coccolithophore blooms, as these blooms lead to a decrease in nutrient flow to lower levels of the ocean.

The most abundant species of coccolithophore, Emiliania huxleyi, belongs to the order Isochrysidales and family Noëlaerhabdaceae. It is found in temperate, subtropical, and tropical oceans. This makes E. huxleyi an important part of the planktonic base of a large proportion of marine food webs. It is also the fastest growing coccolithophore in laboratory cultures. It is studied for the extensive blooms it forms in nutrient depleted waters after the reformation of the summer thermocline. and for its production of molecules known as alkenones that are commonly used by earth scientists as a means to estimate past sea surface temperatures.

Overview
Coccolithophores (or coccolithophorids, from the adjective ) form a group of about 200 phytoplankton species. They belong either to the kingdom Protista, according to Robert Whittaker's Five kingdom classification, or clade Hacrobia, according to the newer biological classification system. Within the Hacrobia, the coccolithophores are in the phylum or division Haptophyta, class Prymnesiophyceae (or Coccolithophyceae). Coccolithophores are distinguished by special calcium carbonate plates (or scales) of uncertain function called coccoliths, which are also important microfossils. However, there are Prymnesiophyceae species lacking coccoliths (e.g. in genus Prymnesium), so not every member of Prymnesiophyceae is a coccolithophore.

Coccolithophores are single-celled phytoplankton that produce small calcium carbonate (CaCO3) scales (coccoliths) which cover the cell surface in the form of a spherical coating, called a coccosphere. They have been an integral part of marine plankton communities since the Jurassic. Today, coccolithophores contribute ~1–10% to inorganic carbon fixation (calcification) to total carbon fixation (calcification plus photosynthesis) in the surface ocean and ~50% to pelagic CaCO3 sediments. Their calcareous shell increases the sinking velocity of photosynthetically fixed into the deep ocean by ballasting organic matter. At the same time, the biogenic precipitation of calcium carbonate during coccolith formation reduces the total alkalinity of seawater and releases. Thus, coccolithophores play an important role in the marine carbon cycle by influencing the efficiency of the biological carbon pump and the oceanic uptake of atmospheric.

As of 2021, it is not known why coccolithophores calcify and how their ability to produce coccoliths is associated with their ecological success. The most plausible benefit of having a coccosphere seems to be a protection against predators or viruses. Viral infection is an important cause of phytoplankton death in the oceans, and it has recently been shown that calcification can influence the interaction between a coccolithophore and its virus. The major predators of marine phytoplankton are microzooplankton like ciliates and dinoflagellates. These are estimated to consume about two-thirds of the primary production in the ocean and microzooplankton can exert a strong grazing pressure on coccolithophore populations. Although calcification does not prevent predation, it has been argued that the coccosphere reduces the grazing efficiency by making it more difficult for the predator to utilise the organic content of coccolithophores. Heterotrophic protists are able to selectively choose prey on the basis of its size or shape and through chemical signals and may thus favor other prey that is available and not protected by coccoliths.

Structure

Coccolithophores are spherical cells about 5–100 micrometres across, enclosed by calcareous plates called coccoliths, which are about 2–25 micrometres across. Each cell contains two brown chloroplasts which surround the nucleus.

Enclosed in each coccosphere is a single cell with membrane bound organelles. Two large chloroplasts with brown pigment are located on either side of the cell and surround the nucleus, mitochondria, golgi apparatus, endoplasmic reticulum, and other organelles. Each cell also has two flagellar structures, which are involved not only in motility, but also in mitosis and formation of the cytoskeleton. In some species, a functional or vestigial haptonema is also present. This structure, which is unique to haptophytes, coils and uncoils in response to environmental stimuli. Although poorly understood, it has been proposed to be involved in prey capture.

Life history strategy

The complex life cycle of coccolithophores is known as a haplodiplontic life cycle, and is characterized by an alternation of both asexual and sexual phases. The asexual phase is known as the haploid phase, while the sexual phase is known as the diploid phase. During the haploid phase, coccolithophores produce haploid cells through mitosis. These haploid cells can then divide further through mitosis or undergo sexual reproduction with other haploid cells. The resulting diploid cell goes through meiosis to produce haploid cells again, starting the cycle over. With coccolithophores, asexual reproduction by mitosis is possible in both phases of the life cycle, which is a contrast with most other organisms that have alternating life cycles. Both abiotic and biotic factors may affect the frequency with which each phase occurs.

Coccolithophores reproduce asexually through binary fission. In this process the coccoliths from the parent cell are divided between the two daughter cells. There have been suggestions stating the possible presence of a sexual reproduction process due to the diploid stages of the coccolithophores, but this process has never been observed.

K or r- selected strategies of coccolithophores depend on their life cycle stage. When coccolithophores are diploid, they are r-selected. In this phase they tolerate a wider range of nutrient compositions. When they are haploid they are K- selected and are often more competitive in stable low nutrient environments. Most coccolithophores are K strategist and are usually found on nutrient-poor surface waters. They are poor competitors when compared to other phytoplankton and thrive in habitats where other phytoplankton would not survive. These two stages in the life cycle of coccolithophores occur seasonally, where more nutrition is available in warmer seasons and less is available in cooler seasons. This type of life cycle is known as a complex heteromorphic life cycle.

Global distribution

Coccolithophores occur throughout the world's oceans. Their distribution varies vertically by stratified layers in the ocean and geographically by different temporal zones. While most modern coccolithophores can be located in their associated stratified oligotrophic conditions, the most abundant areas of coccolithophores where there is the highest species diversity are located in subtropical zones with a temperate climate. While water temperature and the amount of light intensity entering the water's surface are the more influential factors in determining where species are located, the ocean currents also can determine the location where certain species of coccolithophores are found.

Although motility and colony formation vary according to the life cycle of different coccolithophore species, there is often alternation between a motile, haploid phase, and a non-motile diploid phase. In both phases, the organism's dispersal is largely due to ocean currents and circulation patterns.

Within the Pacific Ocean, approximately 90 species have been identified with six separate zones relating to different Pacific currents that contain unique groupings of different species of coccolithophores. The highest diversity of coccolithophores in the Pacific Ocean was in an area of the ocean considered the Central North Zone which is an area between 30 oN and 5 oN, composed of the North Equatorial Current and the Equatorial Countercurrent. These two currents move in opposite directions, east and west, allowing for a strong mixing of waters and allowing a large variety of species to populate the area.

In the Atlantic Ocean, the most abundant species are E. huxleyi and Florisphaera profunda with smaller concentrations of the species Umbellosphaera irregularis, Umbellosphaera tenuis and different species of Gephyrocapsa. Deep-dwelling coccolithophore species abundance is greatly affected by nutricline and thermocline depths. These coccolithophores increase in abundance when the nutricline and thermocline are deep and decrease when they are shallow.

The complete distribution of coccolithophores is currently not known and some regions, such as the Indian Ocean, are not as well studied as other locations in the Pacific and Atlantic Oceans. It is also very hard to explain distributions due to multiple constantly changing factors involving the ocean's properties, such as coastal and equatorial upwelling, frontal systems, benthic environments, unique oceanic topography, and pockets of isolated high or low water temperatures.

The upper photic zone is low in nutrient concentration, high in light intensity and penetration, and usually higher in temperature. The lower photic zone is high in nutrient concentration, low in light intensity and penetration and relatively cool. The middle photic zone is an area that contains the same values in between that of the lower and upper photic zones.

Great Calcite Belt

The Great Calcite Belt of the Southern Ocean is a region of elevated summertime upper ocean calcite concentration derived from coccolithophores, despite the region being known for its diatom predominance. The overlap of two major phytoplankton groups, coccolithophores and diatoms, in the dynamic frontal systems characteristic of this region provides an ideal setting to study environmental influences on the distribution of different species within these taxonomic groups.

The Great Calcite Belt, defined as an elevated particulate inorganic carbon (PIC) feature occurring alongside seasonally elevated chlorophyll a in austral spring and summer in the Southern Ocean, plays an important role in climate fluctuations, accounting for over 60% of the Southern Ocean area (30–60° S). The region between 30° and 50° S has the highest uptake of anthropogenic carbon dioxide (CO2) alongside the North Atlantic and North Pacific oceans.

Effect of global climate change on distribution
Recent studies show that climate change has direct and indirect impacts on Coccolithophore distribution and productivity. They will inevitably be affected by the increasing temperatures and thermal stratification of the top layer of the ocean, since these are prime controls on their ecology, although it is not clear whether global warming would result in net increase or decrease of coccolithophores. As they are calcifying organisms, it has been suggested that ocean acidification due to increasing carbon dioxide could severely affect coccolithophores. Recent increases have seen a sharp increase in the population of coccolithophores.

Role in the food web

Coccolithophores are one of the more abundant primary producers in the ocean. As such, they are a large contributor to the primary productivity of the tropical and subtropical oceans, however, exactly how much has yet to have been recorded.

Dependence on nutrients
The ratio between the concentrations of nitrogen, phosphorus and silicate in particular areas of the ocean dictates competitive dominance within phytoplankton communities. Each ratio essentially tips the odds in favor of either diatoms or other groups of phytoplankton, such as coccolithophores. A low silicate to nitrogen and phosphorus ratio allows coccolithophores to outcompete other phytoplankton species; however, when silicate to phosphorus to nitrogen ratios are high coccolithophores are outcompeted by diatoms. The increase in agricultural processes lead to eutrophication of waters and thus, coccolithophore blooms in these high nitrogen and phosphorus, low silicate environments.

Impact on water column productivity
The calcite in calcium carbonate allows coccoliths to scatter more light than they absorb. This has two important consequences: 1) Surface waters become brighter, meaning they have a higher albedo, and 2) there is induced photoinhibition, meaning photosythetic production is diminished due to an excess of light. In case 1), a high concentration of coccoliths leads to a simultaneous increase in surface water temperature and decrease in the temperature of deeper waters. This results in more stratification in the water column and a decrease in the vertical mixing of nutrients. However, a 2012 study estimated that the overall effect of coccolithophores on the increase in radiative forcing of the ocean is less than that from anthropogenic factors. Therefore, the overall result of large blooms of coccolithophores is a decrease in water column productivity, rather than a contribution to global warming.

Predator-prey interactions
Their predators include the common predators of all phytoplankton including small fish, zooplankton, and shellfish larvae. Viruses specific to this species have been isolated from several locations worldwide and appear to play a major role in spring bloom dynamics.

Toxicity
No environmental evidence of coccolithophore toxicity has been reported, but they belong to the class Prymnesiophyceae which contain orders with toxic species. Toxic species have been found in the genera Prymnesium Massart and Chrysochromulina Lackey. Members of the genus Prymnesium have been found to produce haemolytic compounds, the agent responsible for toxicity. Some of these toxic species are responsible for large fish kills and can be accumulated in organisms such as shellfish; transferring it through the food chain. In laboratory tests for toxicity members of the oceanic coccolithophore genera Emiliania, Gephyrocapsa, Calcidiscus and Coccolithus were shown to be non-toxic as were species of the coastal genus Hymenomonas, however several species of Pleurochrysis and Jomonlithus, both coastal genera were toxic to Artemia.

Community interactions
Coccolithophorids are predominantly found as single, free-floating haploid or diploid cells.

Competition
Most phytoplankton need sunlight and nutrients from the ocean to survive, so they thrive in areas with large inputs of nutrient rich water upwelling from the lower levels of the ocean. Most coccolithophores require sunlight only for energy production, and have a higher ratio of nitrate uptake over ammonium uptake (nitrogen is required for growth and can be used directly from nitrate but not ammonium). Because of this they thrive in still, nutrient-poor environments where other phytoplankton are starving. Trade-offs associated with these faster growth rates include a smaller cell radius and lower cell volume than other types of phytoplankton.

Viral infection and coevolution
Giant DNA-containing viruses are known to lytically infect coccolithophores, particularly E. huxleyi. These viruses, known as E. huxleyi viruses (EhVs), appear to infect the coccosphere coated diploid phase of the life cycle almost exclusively. It has been proposed that as the haploid organism is not infected and therefore not affected by the virus, the co-evolutionary "arms race" between coccolithophores and these viruses does not follow the classic Red Queen evolutionary framework, but instead a "Cheshire Cat" ecological dynamic. More recent work has suggested that viral synthesis of sphingolipids and induction of programmed cell death provides a more direct link to study a Red Queen-like coevolutionary arms race at least between the coccolithoviruses and diploid organism.

Evolution and diversity
Coccolithophores are members of the clade Haptophyta, which is a sister clade to Centrohelida, which are both in Haptista. The oldest known coccolithophores are known from the Late Triassic, around the Norian-Rhaetian boundary. Diversity steadily increased over the course of the Mesozoic, reaching its apex during the Late Cretaceous. However, there was a sharp drop during the Cretaceous-Paleogene extinction event, when more than 90% of coccolithophore species became extinct. Coccoliths reached another, lower apex of diversity during the Paleocene-Eocene thermal maximum, but have subsequently declined since the Oligocene due to decreasing global temperatures, with species that produced large and heavily calcified coccoliths most heavily affected.

Coccolithophore shells

 * Exoskeleton: coccospheres and coccoliths

Each coccolithophore encloses itself in a protective shell of coccoliths, calcified scales which make up its exoskeleton or coccosphere. The coccoliths are created inside the coccolithophore cell and while some species maintain a single layer throughout life only producing new coccoliths as the cell grows, others continually produce and shed coccoliths.

Composition
The primary constituent of coccoliths is calcium carbonate, or chalk. Calcium carbonate is transparent, so the organisms' photosynthetic activity is not compromised by encapsulation in a coccosphere.

Formation
Coccoliths are produced by a biomineralization process known as coccolithogenesis. Generally, calcification of coccoliths occurs in the presence of light, and these scales are produced much more during the exponential phase of growth than the stationary phase. Although not yet entirely understood, the biomineralization process is tightly regulated by calcium signaling. Calcite formation begins in the golgi complex where protein templates nucleate the formation of CaCO3 crystals and complex acidic polysaccharides control the shape and growth of these crystals. As each scale is produced, it is exported in a Golgi-derived vesicle and added to the inner surface of the coccosphere. This means that the most recently produced coccoliths may lie beneath older coccoliths. Depending upon the phytoplankton's stage in the life cycle, two different types of coccoliths may be formed. Holococcoliths are produced only in the haploid phase, lack radial symmetry, and are composed of anywhere from hundreds to thousands of similar minute (ca 0.1 μm) rhombic calcite crystals. These crystals are thought to form at least partially outside the cell. Heterococcoliths occur only in the diploid phase, have radial symmetry, and are composed of relatively few complex crystal units (fewer than 100). Although they are rare, combination coccospheres, which contain both holococcoliths and heterococcoliths, have been observed in the plankton recording coccolithophore life cycle transitions. Finally, the coccospheres of some species are highly modified with various appendages made of specialized coccoliths.

Function
While the exact function of the coccosphere is unclear, many potential functions have been proposed. Most obviously coccoliths may protect the phytoplankton from predators. It also appears that it helps them to create a more stable pH. During photosynthesis carbon dioxide is removed from the water, making it more basic. Also calcification removes carbon dioxide, but chemistry behind it leads to the opposite pH reaction; it makes the water more acidic. The combination of photosynthesis and calcification therefore even out each other regarding pH changes. In addition, these exoskeletons may confer an advantage in energy production, as coccolithogenesis seems highly coupled with photosynthesis. Organic precipitation of calcium carbonate from bicarbonate solution produces free carbon dioxide directly within the cellular body of the alga, this additional source of gas is then available to the Coccolithophore for photosynthesis. It has been suggested that they may provide a cell-wall like barrier to isolate intracellular chemistry from the marine environment. More specific, defensive properties of coccoliths may include protection from osmotic changes, chemical or mechanical shock, and short-wavelength light. It has also been proposed that the added weight of multiple layers of coccoliths allows the organism to sink to lower, more nutrient rich layers of the water and conversely, that coccoliths add buoyancy, stopping the cell from sinking to dangerous depths. Coccolith appendages have also been proposed to serve several functions, such as inhibiting grazing by zooplankton.

Uses
Coccoliths are the main component of the Chalk, a Late Cretaceous rock formation which outcrops widely in southern England and forms the White Cliffs of Dover, and of other similar rocks in many other parts of the world. At the present day sedimented coccoliths are a major component of the calcareous oozes that cover up to 35% of the ocean floor and is kilometres thick in places. Because of their abundance and wide geographic ranges, the coccoliths which make up the layers of this ooze and the chalky sediment formed as it is compacted serve as valuable microfossils.

Calcification, the biological production of calcium carbonate (CaCO3), is a key process in the marine carbon cycle. Coccolithophores are the major planktonic group responsible for pelagic CaCO3 production. The diagram on the right shows the energetic costs of coccolithophore calcification:

 * (A) Transport processes include the transport into the cell from the surrounding seawater of primary calcification substrates Ca2+ and HCO3− (black arrows) and the removal of the end product H+ from the cell (gray arrow). The transport of Ca2+ through the cytoplasm to the CV is the dominant cost associated with calcification.

 * (B) Metabolic processes include the synthesis of CAPs (gray rectangles) by the Golgi complex (white rectangles) that regulate the nucleation and geometry of CaCO3 crystals. The completed coccolith (gray plate) is a complex structure of intricately arranged CAPs and CaCO3 crystals.

 * (C) Mechanical and structural processes account for the secretion of the completed coccoliths that are transported from their original position adjacent to the nucleus to the cell periphery, where they are transferred to the surface of the cell. The costs associated with these processes are likely to be comparable to organic-scale exocytosis in noncalcifying haptophyte algae.

The diagram on the left shows the benefits of coccolithophore calcification. (A) Accelerated photosynthesis includes CCM (1) and enhanced light uptake via scattering of scarce photons for deep-dwelling species (2). (B) Protection from photodamage includes sunshade protection from ultraviolet (UV) light and photosynthetic active radiation (PAR) (1) and energy dissipation under high-light conditions (2). (C) Armor protection includes protection against viral/bacterial infections (1) and grazing by selective (2) and nonselective (3) grazers.

The degree by which calcification can adapt to ocean acidification is presently unknown. Cell physiological examinations found the essential H+ efflux (stemming from the use of HCO3− for intra-cellular calcification) to become more costly with ongoing ocean acidification as the electrochemical H+ inside-out gradient is reduced and passive proton outflow impeded. Adapted cells would have to activate proton channels more frequently, adjust their membrane potential, and/or lower their internal pH. Reduced intra-cellular pH would severely affect the entire cellular machinery and require other processes (e.g. photosynthesis) to co-adapt in order to keep H+ efflux alive. The obligatory H+ efflux associated with calcification may therefore pose a fundamental constraint on adaptation which may potentially explain why "calcification crisis" were possible during long-lasting (thousands of years) CO2 perturbation events even though evolutionary adaption to changing carbonate chemistry conditions is possible within one year. Unraveling these fundamental constraints and the limits of adaptation should be a focus in future coccolithophore studies because knowing them is the key information required to understand to what extent the calcification response to carbonate chemistry perturbations can be compensated by evolution.

Silicate- or cellulose-armored functional groups such as diatoms and dinoflagellates do not need to sustain the calcification-related H+ efflux. Thus, they probably do not need to adapt in order to keep costs for the production of structural elements low. On the contrary, dinoflagellates (except for calcifying species; with generally inefficient CO2-fixing RuBisCO enzymes may even profit from chemical changes since photosynthetic carbon fixation as their source of structural elements in the form of cellulose should be facilitated by the ocean acidification-associated CO2 fertilization. Under the assumption that any form of shell/exoskeleton protects phytoplankton against predation non-calcareous armors may be the preferable solution to realize protection in a future ocean.

The diagram on the right is a representation of how the comparative energetic effort for armor construction in diatoms, dinoflagellates and coccolithophores appear to operate. The frustule (diatom shell) seems to be the most inexpensive armor under all circumstances because diatoms typically outcompete all other groups when silicate is available. The coccosphere is relatively inexpensive under sufficient [CO2], high [HCO3−], and low [H+] because the substrate is saturating and protons are easily released into seawater. In contrast, the construction of thecal elements, which are organic (cellulose) plates that constitute the dinoflagellate shell, should rather be favored at high H+ concentrations because these usually coincide with high [CO2]. Under these conditions dinoflagellates could down-regulate the energy-consuming operation of carbon concentrating mechanisms to fuel the production of organic source material for their shell. Therefore, a shift in carbonate chemistry conditions toward high [CO2] may promote their competitiveness relative to coccolithophores. However, such a hypothetical gain in competitiveness due to altered carbonate chemistry conditions would not automatically lead to dinoflagellate dominance because a huge number of factors other than carbonate chemistry have an influence on species composition as well.

Defence against predation
Currently, the evidence supporting or refuting a protective function of the coccosphere against predation is limited. Some researchers found that overall microzooplankton predation rates were reduced during blooms of the coccolithophore Emiliania huxleyi, while others found high microzooplankton grazing rates on natural coccolithophore communities. In 2020, researchers found that in situ ingestion rates of microzooplankton on E. huxleyi did not differ significantly from those on similar sized non-calcifying phytoplankton. In laboratory experiments the heterotrophic dinoflagellate Oxyrrhis marina preferred calcified over non-calcified cells of E. huxleyi, which was hypothesised to be due to size selective feeding behaviour, since calcified cells are larger than non-calcified E. huxleyi. In 2015, Harvey et al. investigated predation by the dinoflagellate O. marina on different genotypes of non-calcifying E. huxleyi as well as calcified strains that differed in the degree of calcification. They found that the ingestion rate of O. marina was dependent on the genotype of E. huxleyi that was offered, rather than on their degree of calcification. In the same study, however, the authors found that predators which preyed on non-calcifying genotypes grew faster than those fed with calcified cells. In 2018, Strom et al. compared predation rates of the dinoflagellate Amphidinium longum on calcified relative to naked E. huxleyi prey and found no evidence that the coccosphere prevents ingestion by the grazer. Instead, ingestion rates were dependent on the offered genotype of E. huxleyi. Altogether, these two studies suggest that the genotype has a strong influence on ingestion by the microzooplankton species, but if and how calcification protects coccolithophores from microzooplankton predation could not be fully clarified.

Impact on the carbon cycle
Coccolithophores have both long and short term effects on the carbon cycle. The production of coccoliths requires the uptake of dissolved inorganic carbon and calcium. Calcium carbonate and carbon dioxide are produced from calcium and bicarbonate by the following chemical reaction:
 * Ca(2+) + 2HCO3− <-> CaCO3 + CO2 + H2O

Because coccolithophores are photosynthetic organisms, they are able to use some of the released in the calcification reaction for photosynthesis.

However, the production of calcium carbonate drives surface alkalinity down, and in conditions of low alkalinity the is instead released back into the atmosphere. As a result of this, researchers have postulated that large blooms of coccolithophores may contribute to global warming in the short term. A more widely accepted idea, however, is that over the long term coccolithophores contribute to an overall decrease in atmospheric concentrations. During calcification two carbon atoms are taken up and one of them becomes trapped as calcium carbonate. This calcium carbonate sinks to the bottom of the ocean in the form of coccoliths and becomes part of sediment; thus, coccolithophores provide a sink for emitted carbon, mediating the effects of greenhouse gas emissions.

Evolutionary responses to ocean acidification
Research also suggests that ocean acidification due to increasing concentrations of in the atmosphere may affect the calcification machinery of coccolithophores. This may not only affect immediate events such as increases in population or coccolith production, but also may induce evolutionary adaptation of coccolithophore species over longer periods of time. For example, coccolithophores use H+ ion channels in to constantly pump H+ ions out of the cell during coccolith production. This allows them to avoid acidosis, as coccolith production would otherwise produce a toxic excess of H+ ions. When the function of these ion channels is disrupted, the coccolithophores stop the calcification process to avoid acidosis, thus forming a feedback loop. Low ocean alkalinity, impairs ion channel function and therefore places evolutionary selective pressure on coccolithophores and makes them (and other ocean calcifiers) vulnerable to ocean acidification. In 2008, field evidence indicating an increase in calcification of newly formed ocean sediments containing coccolithophores bolstered the first ever experimental data showing that an increase in ocean concentration results in an increase in calcification of these organisms. Decreasing coccolith mass is related to both the increasing concentrations of and decreasing concentrations of CO3(2−) in the world's oceans. This lower calcification is assumed to put coccolithophores at ecological disadvantage. Some species like Calcidiscus leptoporus, however, are not affected in this way, while the most abundant coccolithophore species, E. huxleyi might be (study results are mixed). Also, highly calcified coccolithophorids have been found in conditions of low CaCO3 saturation contrary to predictions. Understanding the effects of increasing ocean acidification on coccolithophore species is absolutely essential to predicting the future chemical composition of the ocean, particularly its carbonate chemistry. Viable conservation and management measures will come from future research in this area. Groups like the European-based CALMARO are monitoring the responses of coccolithophore populations to varying pH's and working to determine environmentally sound measures of control.

Impact on microfossil record
Coccolith fossils are prominent and valuable calcareous microfossils. They are the largest global source of biogenic calcium carbonate, and significantly contribute to the global carbon cycle. They are the main constituent of chalk deposits such as the white cliffs of Dover.

Of particular interest are fossils dating back to the Palaeocene-Eocene Thermal Maximum 55 million years ago. This period is thought to correspond most directly to the current levels of in the ocean. Finally, field evidence of coccolithophore fossils in rock were used to show that the deep-sea fossil record bears a rock record bias similar to the one that is widely accepted to affect the land-based fossil record.

Impact on the oceans
The coccolithophorids help in regulating the temperature of the oceans. They thrive in warm seas and release dimethyl sulfide (DMS) into the air whose nuclei help to produce thicker clouds to block the sun. When the oceans cool, the number of coccolithophorids decrease and the amount of clouds also decrease. When there are fewer clouds blocking the sun, the temperature also rises. This, therefore, maintains the balance and equilibrium of nature.

title: Coccolithophore


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Condensation

Condensation is   the change of the state of matter from the gas phase into the liquid phase, and is the reverse of vaporization. The word most often refers to the water cycle. It can also be defined as the change in the state of water vapor to liquid water when in contact with a liquid or solid surface or cloud condensation nuclei within the atmosphere. When the transition happens from the gaseous phase into the solid phase directly, the change is called deposition.

Initiation
Condensation is initiated by the formation of atomic/molecular clusters of that species within its gaseous volume—like rain drop or snow flake formation within clouds—or at the contact between such gaseous phase and a liquid or solid surface. In clouds, this can be catalyzed by water-nucleating proteins, produced by atmospheric microbes, which are capable of binding gaseous or liquid water molecules.

Reversibility scenarios
A few distinct reversibility scenarios emerge here with respect to the nature of the surface.
 * absorption into the surface of a liquid (either of the same substance or one of its solvents)—is reversible as evaporation.
 * adsorption (as dew droplets) onto solid surface at pressures and temperatures higher than the species' triple point—also reversible as evaporation.
 * adsorption onto solid surface (as supplemental layers of solid) at pressures and temperatures lower than the species' triple point—is reversible as sublimation.

Most common scenarios
Condensation commonly occurs when a vapor is cooled and/or compressed to its saturation limit when the molecular density in the gas phase reaches its maximal threshold. Vapor cooling and compressing equipment that collects condensed liquids is called a "condenser".

Measurement
Psychrometry measures the rates of condensation through evaporation into the air moisture at various atmospheric pressures and temperatures. Water is the product of its vapor condensation—condensation is the process of such phase conversion.

Applications of condensation
Condensation is a crucial component of distillation, an important laboratory and industrial chemistry application.

Because condensation is a naturally occurring phenomenon, it can often be used to generate water in large quantities for human use. Many structures are made solely for the purpose of collecting water from condensation, such as air wells and fog fences. Such systems can often be used to retain soil moisture in areas where active desertification is occurring—so much so that some organizations educate people living in affected areas about water condensers to help them deal effectively with the situation.

It is also a crucial process in forming particle tracks in a cloud chamber. In this case, ions produced by an incident particle act as nucleation centers for the condensation of the vapor producing the visible "cloud" trails.

Commercial applications of condensation, by consumers as well as industry, include power generation, water desalination, thermal management, refrigeration, and air conditioning.

Biological adaptation
Numerous living beings use water made accessible by condensation. A few examples of these are the Australian thorny devil, the darkling beetles of the Namibian coast, and the coast redwoods of the West Coast of the United States.

Condensation in building construction
Condensation in building construction is an unwanted phenomenon as it may cause dampness, mold health issues, wood rot, corrosion, weakening of mortar and masonry walls, and energy penalties due to increased heat transfer. To alleviate these issues, the indoor air humidity needs to be lowered, or air ventilation in the building needs to be improved. This can be done in a number of ways, for example opening windows, turning on extractor fans, using dehumidifiers, drying clothes outside and covering pots and pans whilst cooking. Air conditioning or ventilation systems can be installed that help remove moisture from the air, and move air throughout a building. The amount of water vapor that can be stored in the air can be increased simply by increasing the temperature. However, this can be a double edged sword as most condensation in the home occurs when warm, moisture heavy air comes into contact with a cool surface. As the air is cooled, it can no longer hold as much water vapor. This leads to deposition of water on the cool surface. This is very apparent when central heating is used in combination with single glazed windows in winter.

Interstructure condensation may be caused by thermal bridges, insufficient or lacking insulation, damp proofing or insulated glazing.

license: Creative Commons - Attribution Share-Alike - https://creativecommons.org/licenses/by-sa/4.0/

title: Condensation


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Vandalism?
This might be me but, has this page been vandalised seeing as how I'm looking at offensive/just plain stupid text? Darth Gladius 22:28, 7 June 2007 (UTC)

title: Talk:Movies


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Contrail

Contrails (short for "condensation trails") or vapor trails are line-shaped clouds produced by aircraft engine exhaust or changes in air pressure, typically at aircraft cruising altitudes several kilometres/miles above the Earth's surface. They are composed primarily of water, in the form of ice crystals. The combination of water vapor in aircraft engine exhaust and the low ambient temperatures at high altitudes causes the trails' formation. Impurities in the engine exhaust from the fuel, including sulfur compounds (0.05% by weight in jet fuel) provide some of the particles that serve as cloud condensation nuclei for water droplet growth in the exhaust. If water droplets form, they can freeze to form ice particles that compose a contrail. Their formation can also be triggered by changes in air pressure in wingtip vortices, or in the air over the entire wing surface. Contrails, and other clouds caused directly by human activity, are called homogenitus.

Depending on the temperature and humidity at the altitude where the contrails form, they may be visible for only a few seconds or minutes. Persistent contrails are of particular interest to scientists. The resulting cloud forms are formally described as homomutatus, and may resemble cirrus, cirrocumulus, or cirrostratus, and are sometimes called cirrus aviaticus. Some persistent spreading contrails contribute to climate change.

Condensation trails as a result of engine exhaust

Engine exhaust is predominantly made up of water and carbon dioxide, the combustion products of hydrocarbon fuels. Many other chemical byproducts of incomplete hydrocarbon fuel combustion, including volatile organic compounds, inorganic gases, polycyclic aromatic hydrocarbons, oxygenated organics, alcohols, ozone and particles of soot have been observed at lower concentrations. The exact quality is a function of engine type and basic combustion engine function, with up to 30% of aircraft exhaust being unburned fuel. (Micron-sized metallic particles resulting from engine wear have also been detected.) At high altitudes as this water vapor emerges into a cold environment, the localized increase in water vapor can raise the relative humidity of the air past saturation point. The vapor then condenses into tiny water droplets which freeze if the temperature is low enough. These millions of tiny water droplets and/or ice crystals form the contrails. The time taken for the vapor to cool enough to condense accounts for the contrail forming some distance behind the aircraft. At high altitudes, supercooled water vapor requires a trigger to encourage deposition or condensation. The exhaust particles in the aircraft's exhaust act as this trigger, causing the trapped vapor to condense rapidly. Exhaust contrails usually form at high altitudes; usually above 8000 m, where the air temperature is below -36.5 C. They can also form closer to the ground when the air is cold and moist.

A 2013–2014 study jointly supported by NASA, the German aerospace center DLR, and Canada's National Research Council NRC, determined that biofuels could reduce contrail generation. This reduction was explained by demonstrating that biofuels produce fewer soot particles, which are the nuclei around which the ice crystals form. The tests were performed by flying a DC-8 at cruising altitude with a sample-gathering aircraft flying in trail. In these samples, the contrail-producing soot particle count was reduced by 50 to 70 percent, using a 50% blend of conventional Jet A1 fuel and HEFA (hydroprocessed esters and fatty acids) biofuel produced from camelina.

Condensation from decreases in pressure

As a wing generates lift, it causes a vortex to form at the wingtip, and at the tip of the flap when deployed (wingtips and flap boundaries represent discontinuities in airflow). These wingtip vortices persist in the atmosphere long after the aircraft has passed. The reduction in pressure and temperature across each vortex can cause water to condense and make the cores of the wingtip vortices visible; this effect is more common on humid days. Wingtip vortices can sometimes be seen behind the wing flaps of airliners during takeoff and landing, and during Space Shuttle landings.

The visible cores of wingtip vortices contrast with the other major type of contrails which are caused by the combustion of fuel. Contrails produced from jet engine exhaust are seen at high altitude, directly behind each engine. By contrast, the visible cores of wingtip vortices are usually seen only at low altitude where the aircraft is travelling slowly after takeoff or before landing, and where the ambient humidity is higher; they trail behind the wingtips and wing flaps rather than behind the engines.

At high-thrust settings the fan blades at the intake of a turbofan engine reach transonic speeds, causing a sudden drop in air pressure. This creates the condensation fog (inside the intake) which is often observed by air travelers during takeoff.

The tips of rotating surfaces (such as propellers and rotors) sometimes produce visible contrails.

In firearms, a vapor trail is sometimes observed when firing under rare conditions, due to condensation induced by changes in air pressure around the bullet. A vapor trail from a bullet is observable from any direction. Vapor trail should not be confused with bullet trace, a refractive effect due to changes in air pressure as the bullet travels, which is a much more common phenomenon (and is usually only observable directly from behind the shooter).

Impacts on climate
It is considered that the largest contribution of aviation to climate change comes from contrails. In general, aircraft contrails trap outgoing longwave radiation emitted by the Earth and atmosphere more than they reflect incoming solar radiation, resulting in a net increase in radiative forcing. In 1992, this warming effect was estimated between 3.5 mW/m2 and 17 mW/m2. In 2009, its 2005 value was estimated at 12 mW/m2, based on the reanalysis data, climate models, and radiative transfer codes; with an uncertainty range of 5 to 26 mW/m2, and with a low level of scientific understanding. Contrail cirrus may be air traffic's largest radiative forcing component, larger than all accumulated from aviation, and could triple from a 2006 baseline to 160–180 mW/m2 by 2050 without intervention. For comparison, the total radiative forcing from human activities amounted to 2.72 W/m2 (with a range between 1.96 and 3.48W/m2) in 2019, and the increase from 2011 to 2019 alone amounted to 0.34W/m2. Contrail effects differ a lot depending on when they are formed, as they decrease the daytime temperature and increase the nighttime temperature, reducing their difference. In 2006, it was estimated that night flights contribute 60 to 80% of contrail radiative forcing while accounting for 25% of daily air traffic, and winter flights contribute half of the annual mean radiative forcing while accounting for 22% of annual air traffic.

Starting from the 1990s, it was suggested that contrails during daytime have a strong cooling effect, and when combined with the warming from night-time flights, this would lead to a substantial diurnal temperature variation (the difference in the day's highs and lows at a fixed station). When no commercial aircraft flew across the USA following the September 11 attacks, the diurnal temperature variation was widened by 1.1 C-change. Measured across 4,000 weather stations in the continental United States, this increase was the largest recorded in 30 years. Without contrails, the local diurnal temperature range was 1 °C-change higher than immediately before. In the southern US, the difference was diminished by about 6 F-change, and by 5 F-change in the US midwest. However, follow-up studies found that a natural change in cloud cover can more than explain these findings. The authors of a 2008 study wrote, "The variations in high cloud cover, including contrails and contrail-induced cirrus clouds, contribute weakly to the changes in the diurnal temperature range, which is governed primarily by lower altitude clouds, winds, and humidity." In 2011, a study of British meteorological records taken during World War II identified one event where the temperature was 0.8 °C-change higher than the day's average near airbases used by USAAF strategic bombers after they flew in a formation. However, its authors cautioned that this was a single event, making it difficult to draw firm conclusions from it. Then, the global response to the 2020 coronavirus pandemic led to a reduction in global air traffic of nearly 70% relative to 2019. Thus, it provided an extended opportunity to study the impact of contrails on regional and global temperature. Multiple studies found "no significant response of diurnal surface air temperature range" as the result of contrail changes, and either "no net significant global ERF" (effective radiative forcing) or a very small warming effect.

An EU project launched in 2020 aims to assess the feasibility of minimising contrail effects by the operational choices in making flight plans. Other similar projects include ContrailNet from Eurocontrol,   Reviate, and the Ciconia project, as well as Google's 'project contrail'.

Head-on contrails
A contrail from an airplane flying towards the observer can appear to be generated by an object moving vertically. On 8 November 2010 in the US state of California, a contrail of this type gained media attention as a "mystery missile" that could not be explained by U.S. military and aviation authorities, and its explanation as a contrail  took more than 24 hours to become accepted by U.S. media and military institutions.

Distrails

Where an aircraft passes through a cloud, it can disperse the cloud in its path. This is known as a distrail (short for "dissipation trail"). The plane's warm engine exhaust and enhanced vertical mixing in the aircraft's wake can cause existing cloud droplets to evaporate. If the cloud is sufficiently thin, such processes can yield a cloud-free corridor in an otherwise solid cloud layer. An early satellite observation of distrails that most likely were elongated, aircraft-induced fallstreak holes appeared in Corfidi and Brandli (1986).

Clouds form when invisible water vapor ( in gas phase) condenses into microscopic water droplets ( in liquid phase) or into microscopic ice crystals ( in solid phase). This may happen when air with a high proportion of gaseous water cools. A distrail forms when the heat of engine exhaust evaporates the liquid water droplets in a cloud, turning them back into invisible, gaseous water vapor. Distrails also may arise as a result of enhanced mixing (entrainment of) drier air immediately above or below a thin cloud layer following passage of an aircraft through the cloud, as shown in the second image below:

title: Contrail


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Convection

Convection is single or multiphase fluid flow that occurs spontaneously due to the combined effects of material property heterogeneity and body forces on a fluid, most commonly density and gravity (see buoyancy). When the cause of the convection is unspecified, convection due to the effects of thermal expansion and buoyancy can be assumed. Convection may also take place in soft solids or mixtures where particles can flow.

Convective flow may be transient (such as when a multiphase mixture of oil and water separates) or steady state (see convection cell). The convection may be due to gravitational, electromagnetic or fictitious body forces. Heat transfer by natural convection plays a role in the structure of Earth's atmosphere, its oceans, and its mantle. Discrete convective cells in the atmosphere can be identified by clouds, with stronger convection resulting in thunderstorms. Natural convection also plays a role in stellar physics. Convection is often categorised or described by the main effect causing the convective flow; for example, thermal convection.

Convection cannot take place in most solids because neither bulk current flows nor significant diffusion of matter can take place. Granular convection is a similar phenomenon in granular material instead of fluids. Advection is fluid motion created by velocity instead of thermal gradients. Convective heat transfer is the intentional use of convection as a method for heat transfer. Convection is a process in which heat is carried from place to place by the bulk movement of a fluid and gases.

History
In the 1830s, in The Bridgewater Treatises, the term convection is attested in a scientific sense. In treatise VIII by William Prout, in the book on chemistry, it says:

[...] This motion of heat takes place in three ways, which a common fire-place very well illustrates. If, for instance, we place a thermometer directly before a fire, it soon begins to rise, indicating an increase of temperature. In this case the heat has made its way through the space between the fire and the thermometer, by the process termed radiation. If we place a second thermometer in contact with any part of the grate, and away from the direct influence of the fire, we shall find that this thermometer also denotes an increase of temperature; but here the heat must have travelled through the metal of the grate, by what is termed conduction. Lastly, a third thermometer placed in the chimney, away from the direct influence of the fire, will also indicate a considerable increase of temperature; in this case a portion of the air, passing through and near the fire, has become heated, and has carried up the chimney the temperature acquired from the fire. There is at present no single term in our language employed to denote this third mode of the propagation of heat; but we venture to propose for that purpose, the term convection, [in footnote: [Latin] Convectio, a carrying or conveying] which not only expresses the leading fact, but also accords very well with the two other terms.

Later, in the same treatise VIII, in the book on meteorology, the concept of convection is also applied to "the process by which heat is communicated through water".

Terminology
Today, the word convection has different but related usages in different scientific or engineering contexts or applications.

In fluid mechanics, convection has a broader sense: it refers to the motion of fluid driven by density (or other property) difference.

In thermodynamics, convection often refers to heat transfer by convection, where the prefixed variant Natural Convection is used to distinguish the fluid mechanics concept of Convection (covered in this article) from convective heat transfer.

Some phenomena which result in an effect superficially similar to that of a convective cell may also be (inaccurately) referred to as a form of convection; for example, thermo-capillary convection and granular convection.

Mechanisms
Convection may happen in fluids at all scales larger than a few atoms. There are a variety of circumstances in which the forces required for convection arise, leading to different types of convection, described below. In broad terms, convection arises because of body forces acting within the fluid, such as gravity.

Natural convection
Natural convection is a flow whose motion is caused by some parts of a fluid being heavier than other parts. In most cases this leads to natural circulation: the ability of a fluid in a system to circulate continuously under gravity, with transfer of heat energy.

The driving force for natural convection is gravity. In a column of fluid, pressure increases with depth from the weight of the overlying fluid. The pressure at the bottom of a submerged object then exceeds that at the top, resulting in a net upward buoyancy force equal to the weight of the displaced fluid. Objects of higher density than that of the displaced fluid then sink. For example, regions of warmer low-density air rise, while those of colder high-density air sink. This creates a circulating flow: convection.

Gravity drives natural convection. Without gravity, convection does not occur, so there is no convection in free-fall (inertial) environments, such as that of the orbiting International Space Station. Natural convection can occur when there are hot and cold regions of either air or water, because both water and air become less dense as they are heated. But, for example, in the world's oceans it also occurs due to salt water being heavier than fresh water, so a layer of salt water on top of a layer of fresher water will also cause convection.

Natural convection has attracted a great deal of attention from researchers because of its presence both in nature and engineering applications. In nature, convection cells formed from air raising above sunlight-warmed land or water are a major feature of all weather systems. Convection is also seen in the rising plume of hot air from fire, plate tectonics, oceanic currents (thermohaline circulation) and sea-wind formation (where upward convection is also modified by Coriolis forces). In engineering applications, convection is commonly visualized in the formation of microstructures during the cooling of molten metals, and fluid flows around shrouded heat-dissipation fins, and solar ponds. A very common industrial application of natural convection is free air cooling without the aid of fans: this can happen on small scales (computer chips) to large scale process equipment.

Natural convection will be more likely and more rapid with a greater variation in density between the two fluids, a larger acceleration due to gravity that drives the convection or a larger distance through the convecting medium. Natural convection will be less likely and less rapid with more rapid diffusion (thereby diffusing away the thermal gradient that is causing the convection) or a more viscous (sticky) fluid.

The onset of natural convection can be determined by the Rayleigh number (Ra).

Differences in buoyancy within a fluid can arise for reasons other than temperature variations, in which case the fluid motion is called gravitational convection (see below). However, all types of buoyant convection, including natural convection, do not occur in microgravity environments. All require the presence of an environment which experiences g-force (proper acceleration).

The difference of density in the fluid is the key driving mechanism. If the differences of density are caused by heat, this force is called as "thermal head" or "thermal driving head." A fluid system designed for natural circulation will have a heat source and a heat sink. Each of these is in contact with some of the fluid in the system, but not all of it. The heat source is positioned lower than the heat sink.

Most fluids expand when heated, becoming less dense, and contract when cooled, becoming denser. At the heat source of a system of natural circulation, the heated fluid becomes lighter than the fluid surrounding it, and thus rises. At the heat sink, the nearby fluid becomes denser as it cools, and is drawn downward by gravity. Together, these effects create a flow of fluid from the heat source to the heat sink and back again.

Gravitational or buoyant convection
Gravitational convection is a type of natural convection induced by buoyancy variations resulting from material properties other than temperature. Typically this is caused by a variable composition of the fluid. If the varying property is a concentration gradient, it is known as solutal convection. For example, gravitational convection can be seen in the diffusion of a source of dry salt downward into wet soil due to the buoyancy of fresh water in saline.

Variable salinity in water and variable water content in air masses are frequent causes of convection in the oceans and atmosphere which do not involve heat, or else involve additional compositional density factors other than the density changes from thermal expansion (see thermohaline circulation). Similarly, variable composition within the Earth's interior which has not yet achieved maximal stability and minimal energy (in other words, with densest parts deepest) continues to cause a fraction of the convection of fluid rock and molten metal within the Earth's interior (see below).

Gravitational convection, like natural thermal convection, also requires a g-force environment in order to occur.

Solid-state convection in ice
Ice convection on Pluto is believed to occur in a soft mixture of nitrogen ice and carbon monoxide ice. It has also been proposed for Europa, and other bodies in the outer Solar System.

Thermomagnetic convection
Thermomagnetic convection can occur when an external magnetic field is imposed on a ferrofluid with varying magnetic susceptibility. In the presence of a temperature gradient this results in a nonuniform magnetic body force, which leads to fluid movement. A ferrofluid is a liquid which becomes strongly magnetized in the presence of a magnetic field.

Combustion
In a zero-gravity environment, there can be no buoyancy forces, and thus no convection possible, so flames in many circumstances without gravity smother in their own waste gases. Thermal expansion and chemical reactions resulting in expansion and contraction gases allows for ventilation of the flame, as waste gases are displaced by cool, fresh, oxygen-rich gas. moves in to take up the low pressure zones created when flame-exhaust water condenses.

Examples and applications
Systems of natural circulation include tornadoes and other weather systems, ocean currents, and household ventilation. Some solar water heaters use natural circulation. The Gulf Stream circulates as a result of the evaporation of water. In this process, the water increases in salinity and density. In the North Atlantic Ocean, the water becomes so dense that it begins to sink down.

Convection occurs on a large scale in atmospheres, oceans, planetary mantles, and it provides the mechanism of heat transfer for a large fraction of the outermost interiors of the Sun and all stars. Fluid movement during convection may be invisibly slow, or it may be obvious and rapid, as in a hurricane. On astronomical scales, convection of gas and dust is thought to occur in the accretion disks of black holes, at speeds which may closely approach that of light.

Demonstration experiments
Thermal convection in liquids can be demonstrated by placing a heat source (for example, a Bunsen burner) at the side of a container with a liquid. Adding a dye to the water (such as food colouring) will enable visualisation of the flow.

Another common experiment to demonstrate thermal convection in liquids involves submerging open containers of hot and cold liquid coloured with dye into a large container of the same liquid without dye at an intermediate temperature (for example, a jar of hot tap water coloured red, a jar of water chilled in a fridge coloured blue, lowered into a clear tank of water at room temperature).

A third approach is to use two identical jars, one filled with hot water dyed one colour, and cold water of another colour. One jar is then temporarily sealed (for example, with a piece of card), inverted and placed on top of the other. When the card is removed, if the jar containing the warmer liquid is placed on top no convection will occur. If the jar containing colder liquid is placed on top, a convection current will form spontaneously.

Convection in gases can be demonstrated using a candle in a sealed space with an inlet and exhaust port. The heat from the candle will cause a strong convection current which can be demonstrated with a flow indicator, such as smoke from another candle, being released near the inlet and exhaust areas respectively.

Convection cells

A convection cell, also known as a Bénard cell, is a characteristic fluid flow pattern in many convection systems. A rising body of fluid typically loses heat because it encounters a colder surface. In liquid, this occurs because it exchanges heat with colder liquid through direct exchange. In the example of the Earth's atmosphere, this occurs because it radiates heat. Because of this heat loss the fluid becomes denser than the fluid underneath it, which is still rising. Since it cannot descend through the rising fluid, it moves to one side. At some distance, its downward force overcomes the rising force beneath it, and the fluid begins to descend. As it descends, it warms again and the cycle repeats itself. Additionally, convection cells can arise due to density variations resulting from differences in the composition of electrolytes.

Atmospheric circulation

Atmospheric circulation is the large-scale movement of air, and is a means by which thermal energy is distributed on the surface of the Earth, together with the much slower (lagged) ocean circulation system. The large-scale structure of the atmospheric circulation varies from year to year, but the basic climatological structure remains fairly constant.

Latitudinal circulation occurs because incident solar radiation per unit area is highest at the heat equator, and decreases as the latitude increases, reaching minima at the poles. It consists of two primary convection cells, the Hadley cell and the polar vortex, with the Hadley cell experiencing stronger convection due to the release of latent heat energy by condensation of water vapor at higher altitudes during cloud formation.

Longitudinal circulation, on the other hand, comes about because the ocean has a higher specific heat capacity than land (and also thermal conductivity, allowing the heat to penetrate further beneath the surface ) and thereby absorbs and releases more heat, but the temperature changes less than land. This brings the sea breeze, air cooled by the water, ashore in the day, and carries the land breeze, air cooled by contact with the ground, out to sea during the night. Longitudinal circulation consists of two cells, the Walker circulation and El Niño / Southern Oscillation.

Weather

Some more localized phenomena than global atmospheric movement are also due to convection, including wind and some of the hydrologic cycle. For example, a foehn wind is a down-slope wind which occurs on the downwind side of a mountain range. It results from the adiabatic warming of air which has dropped most of its moisture on windward slopes. Because of the different adiabatic lapse rates of moist and dry air, the air on the leeward slopes becomes warmer than at the same height on the windward slopes.

A thermal column (or thermal) is a vertical section of rising air in the lower altitudes of the Earth's atmosphere. Thermals are created by the uneven heating of the Earth's surface from solar radiation. The Sun warms the ground, which in turn warms the air directly above it. The warmer air expands, becoming less dense than the surrounding air mass, and creating a thermal low. The mass of lighter air rises, and as it does, it cools by expansion at lower air pressures. It stops rising when it has cooled to the same temperature as the surrounding air. Associated with a thermal is a downward flow surrounding the thermal column. The downward moving exterior is caused by colder air being displaced at the top of the thermal. Another convection-driven weather effect is the sea breeze.

Warm air has a lower density than cool air, so warm air rises within cooler air, similar to hot air balloons. Clouds form as relatively warmer air carrying moisture rises within cooler air. As the moist air rises, it cools, causing some of the water vapor in the rising packet of air to condense. When the moisture condenses, it releases energy known as latent heat of condensation which allows the rising packet of air to cool less than its surrounding air, continuing the cloud's ascension. If enough instability is present in the atmosphere, this process will continue long enough for cumulonimbus clouds to form, which support lightning and thunder. Generally, thunderstorms require three conditions to form: moisture, an unstable airmass, and a lifting force (heat).

All thunderstorms, regardless of type, go through three stages: the developing stage, the mature stage, and the dissipation stage. The average thunderstorm has a 24 km diameter. Depending on the conditions present in the atmosphere, these three stages take an average of 30 minutes to go through.

Oceanic circulation

Solar radiation affects the oceans: warm water from the Equator tends to circulate toward the poles, while cold polar water heads towards the Equator. The surface currents are initially dictated by surface wind conditions. The trade winds blow westward in the tropics, and the westerlies blow eastward at mid-latitudes. This wind pattern applies a stress to the subtropical ocean surface with negative curl across the Northern Hemisphere, and the reverse across the Southern Hemisphere. The resulting Sverdrup transport is equatorward. Because of conservation of potential vorticity caused by the poleward-moving winds on the subtropical ridge's western periphery and the increased relative vorticity of poleward moving water, transport is balanced by a narrow, accelerating poleward current, which flows along the western boundary of the ocean basin, outweighing the effects of friction with the cold western boundary current which originates from high latitudes. The overall process, known as western intensification, causes currents on the western boundary of an ocean basin to be stronger than those on the eastern boundary.

As it travels poleward, warm water transported by strong warm water current undergoes evaporative cooling. The cooling is wind driven: wind moving over water cools the water and also causes evaporation, leaving a saltier brine. In this process, the water becomes saltier and denser. and decreases in temperature. Once sea ice forms, salts are left out of the ice, a process known as brine exclusion. These two processes produce water that is denser and colder. The water across the northern Atlantic Ocean becomes so dense that it begins to sink down through less salty and less dense water. (This open ocean convection is not unlike that of a lava lamp.) This downdraft of heavy, cold and dense water becomes a part of the North Atlantic Deep Water, a south-going stream.

Mantle convection

Mantle convection is the slow creeping motion of Earth's rocky mantle caused by convection currents carrying heat from the interior of the Earth to the surface. It is one of 3 driving forces that causes tectonic plates to move around the Earth's surface.

The Earth's surface is divided into a number of tectonic plates that are continuously being created and consumed at their opposite plate boundaries. Creation (accretion) occurs as mantle is added to the growing edges of a plate. This hot added material cools down by conduction and convection of heat. At the consumption edges of the plate, the material has thermally contracted to become dense, and it sinks under its own weight in the process of subduction at an ocean trench. This subducted material sinks to some depth in the Earth's interior where it is prohibited from sinking further. The subducted oceanic crust triggers volcanism.

Convection within Earth's mantle is the driving force for plate tectonics. Mantle convection is the result of a thermal gradient: the lower mantle is hotter than the upper mantle, and is therefore less dense. This sets up two primary types of instabilities. In the first type, plumes rise from the lower mantle, and corresponding unstable regions of lithosphere drip back into the mantle. In the second type, subducting oceanic plates (which largely constitute the upper thermal boundary layer of the mantle) plunge back into the mantle and move downwards towards the core-mantle boundary. Mantle convection occurs at rates of centimeters per year, and it takes on the order of hundreds of millions of years to complete a cycle of convection.

Neutrino flux measurements from the Earth's core (see kamLAND) show the source of about two-thirds of the heat in the inner core is the radioactive decay of 40K, uranium and thorium. This has allowed plate tectonics on Earth to continue far longer than it would have if it were simply driven by heat left over from Earth's formation; or with heat produced from gravitational potential energy, as a result of physical rearrangement of denser portions of the Earth's interior toward the center of the planet (that is, a type of prolonged falling and settling).

Stack effect
The Stack effect or chimney effect is the movement of air into and out of buildings, chimneys, flue gas stacks, or other containers due to buoyancy. Buoyancy occurs due to a difference in indoor-to-outdoor air density resulting from temperature and moisture differences. The greater the thermal difference and the height of the structure, the greater the buoyancy force, and thus the stack effect. The stack effect helps drive natural ventilation and infiltration. Some cooling towers operate on this principle; similarly the solar updraft tower is a proposed device to generate electricity based on the stack effect.

Stellar physics

The convection zone of a star is the range of radii in which energy is transported outward from the core region primarily by convection rather than radiation. This occurs at radii which are sufficiently opaque that convection is more efficient than radiation at transporting energy.

Granules on the photosphere of the Sun are the visible tops of convection cells in the photosphere, caused by convection of plasma in the photosphere. The rising part of the granules is located in the center where the plasma is hotter. The outer edge of the granules is darker due to the cooler descending plasma. A typical granule has a diameter on the order of 1,000 kilometers and each lasts 8 to 20 minutes before dissipating. Below the photosphere is a layer of much larger "supergranules" up to 30,000 kilometers in diameter, with lifespans of up to 24 hours.

Water convection at freezing temperatures
Water is a fluid that does not obey the Boussinesq approximation. This is because its density varies nonlinearly with temperature, which causes its thermal expansion coefficient to be inconsistent near freezing temperatures. The density of water reaches a maximum at 4 °C and decreases as the temperature deviates. This phenomenon is investigated by experiment and numerical methods. Water is initially stagnant at 10 °C within a square cavity. It is differentially heated between the two vertical walls, where the left and right walls are held at 10 °C and 0 °C, respectively. The density anomaly manifests in its flow pattern. As the water is cooled at the right wall, the density increases, which accelerates the flow downward. As the flow develops and the water cools further, the decrease in density causes a recirculation current at the bottom right corner of the cavity.

Another case of this phenomenon is the event of super-cooling, where the water is cooled to below freezing temperatures but does not immediately begin to freeze. Under the same conditions as before, the flow is developed. Afterward, the temperature of the right wall is decreased to −10 °C. This causes the water at that wall to become supercooled, create a counter-clockwise flow, and initially overpower the warm current. This plume is caused by a delay in the nucleation of the ice. Once ice begins to form, the flow returns to a similar pattern as before and the solidification propagates gradually until the flow is redeveloped.

Nuclear reactors
In a nuclear reactor, natural circulation can be a design criterion. It is achieved by reducing turbulence and friction in the fluid flow (that is, minimizing head loss), and by providing a way to remove any inoperative pumps from the fluid path. Also, the reactor (as the heat source) must be physically lower than the steam generators or turbines (the heat sink). In this way, natural circulation will ensure that the fluid will continue to flow as long as the reactor is hotter than the heat sink, even when power cannot be supplied to the pumps. Notable examples are the S5G and S8G  United States Naval reactors, which were designed to operate at a significant fraction of full power under natural circulation, quieting those propulsion plants. The S6G reactor cannot operate at power under natural circulation, but can use it to maintain emergency cooling while shut down.

By the nature of natural circulation, fluids do not typically move very fast, but this is not necessarily bad, as high flow rates are not essential to safe and effective reactor operation. In modern design nuclear reactors, flow reversal is almost impossible. All nuclear reactors, even ones designed to primarily use natural circulation as the main method of fluid circulation, have pumps that can circulate the fluid in the case that natural circulation is not sufficient.

Mathematical models of convection
A number of dimensionless terms have been derived to describe and predict convection, including the Archimedes number, Grashof number, Richardson number, and the Rayleigh number.

In cases of mixed convection (natural and forced occurring together) one would often like to know how much of the convection is due to external constraints, such as the fluid velocity in the pump, and how much is due to natural convection occurring in the system.

The relative magnitudes of the Grashof number and the square of the Reynolds number determine which form of convection dominates. If $$\rm Gr/Re^2 \gg 1 $$, forced convection may be neglected, whereas if $$\rm Gr/Re^2 \ll 1 $$, natural convection may be neglected. If the ratio, known as the Richardson number, is approximately one, then both forced and natural convection need to be taken into account.

Onset
The onset of natural convection is determined by the Rayleigh number (Ra). This dimensionless number is given by

 * $$\textbf{Ra} = \frac{\Delta\rho g L^3}{D\mu}$$

where

 * $$\Delta \rho$$ is the difference in density between the two parcels of material that are mixing
 * $$g$$ is the local gravitational acceleration
 * $$L$$ is the characteristic length-scale of convection: the depth of the boiling pot, for example
 * $$D$$ is the diffusivity of the characteristic that is causing the convection, and
 * $$\mu$$ is the dynamic viscosity.

Natural convection will be more likely and/or more rapid with a greater variation in density between the two fluids, a larger acceleration due to gravity that drives the convection, and/or a larger distance through the convecting medium. Convection will be less likely and/or less rapid with more rapid diffusion (thereby diffusing away the gradient that is causing the convection) and/or a more viscous (sticky) fluid.

For thermal convection due to heating from below, as described in the boiling pot above, the equation is modified for thermal expansion and thermal diffusivity. Density variations due to thermal expansion are given by:

 * $$\Delta\rho=\rho_0 \beta \Delta T$$

where

 * $$\rho_0$$ is the reference density, typically picked to be the average density of the medium,
 * $$\beta$$ is the coefficient of thermal expansion, and
 * $$\Delta T$$ is the temperature difference across the medium.

The general diffusivity, $$D$$, is redefined as a thermal diffusivity, $$\alpha$$.

 * $$D=\alpha$$

Inserting these substitutions produces a Rayleigh number that can be used to predict thermal convection.

 * $$\textbf{Ra} = \frac{\rho_0 g \beta \Delta T L^3}{\alpha \mu}$$

Turbulence
The tendency of a particular naturally convective system towards turbulence relies on the Grashof number (Gr).

 * $$ Gr= \frac{g \beta \Delta T L^3}{\nu^2} $$

In very sticky, viscous fluids (large ν), fluid motion is restricted, and natural convection will be non-turbulent.

Following the treatment of the previous subsection, the typical fluid velocity is of the order of $$g \Delta \rho L^2 / \mu$$, up to a numerical factor depending on the geometry of the system. Therefore, Grashof number can be thought of as Reynolds number with the velocity of natural convection replacing the velocity in Reynolds number's formula. However In practice, when referring to the Reynolds number, it is understood that one is considering forced convection, and the velocity is taken as the velocity dictated by external constraints (see below).

Behavior
The Grashof number can be formulated for natural convection occurring due to a concentration gradient, sometimes termed thermo-solutal convection. In this case, a concentration of hot fluid diffuses into a cold fluid, in much the same way that ink poured into a container of water diffuses to dye the entire space. Then:

 * $$ Gr= \frac{g \beta \Delta C L^3}{\nu^2} $$

Natural convection is highly dependent on the geometry of the hot surface, various correlations exist in order to determine the heat transfer coefficient. A general correlation that applies for a variety of geometries is

 * $$Nu = \left[Nu_0^\frac{1}{2} + Ra^ \frac{1}{6} \left(\frac {f_4\left(Pr\right)}{300}\right)^\frac{1}{6} \right]^2 $$

The value of f4(Pr) is calculated using the following formula

 * $$f_4(Pr)= \left[1+ \left ( \frac {0.5}{Pr} \right )^\frac{9}{16} \right]^\frac{-16}{9}$$

Nu is the Nusselt number and the values of Nu0 and the characteristic length used to calculate Re are listed below (see also Discussion):

Warning: The values indicated for the Horizontal cylinder are wrong; see discussion.

Natural convection from a vertical plate
One example of natural convection is heat transfer from an isothermal vertical plate immersed in a fluid, causing the fluid to move parallel to the plate. This will occur in any system wherein the density of the moving fluid varies with position. These phenomena will only be of significance when the moving fluid is minimally affected by forced convection.

When considering the flow of fluid is a result of heating, the following correlations can be used, assuming the fluid is an ideal diatomic, has adjacent to a vertical plate at constant temperature and the flow of the fluid is completely laminar.

Num = 0.478(Gr0.25)

Mean Nusselt number = Num = hmL/k

where

 * hm = mean coefficient applicable between the lower edge of the plate and any point in a distance L (W/m2. K)
 * L = height of the vertical surface (m)
 * k = thermal conductivity (W/m. K)

Grashof number = Gr = $$[gL^3(t_s-t_\infty)]/v^2T$$

where

 * g = gravitational acceleration (m/s2)
 * L = distance above the lower edge (m)
 * ts = temperature of the wall (K)
 * t∞ = fluid temperature outside the thermal boundary layer (K)
 * v = kinematic viscosity of the fluid (m2/s)
 * T = absolute temperature (K)

When the flow is turbulent different correlations involving the Rayleigh Number (a function of both the Grashof number and the Prandtl number) must be used.

Note that the above equation differs from the usual expression for Grashof number because the value $$\beta$$ has been replaced by its approximation $$1/T$$, which applies for ideal gases only (a reasonable approximation for air at ambient pressure).

Pattern formation

Convection, especially Rayleigh–Bénard convection, where the convecting fluid is contained by two rigid horizontal plates, is a convenient example of a pattern-forming system.

When heat is fed into the system from one direction (usually below), at small values it merely diffuses (conducts) from below upward, without causing fluid flow. As the heat flow is increased, above a critical value of the Rayleigh number, the system undergoes a bifurcation from the stable conducting state to the convecting state, where bulk motion of the fluid due to heat begins. If fluid parameters other than density do not depend significantly on temperature, the flow profile is symmetric, with the same volume of fluid rising as falling. This is known as Boussinesq convection.

As the temperature difference between the top and bottom of the fluid becomes higher, significant differences in fluid parameters other than density may develop in the fluid due to temperature. An example of such a parameter is viscosity, which may begin to significantly vary horizontally across layers of fluid. This breaks the symmetry of the system, and generally changes the pattern of up- and down-moving fluid from stripes to hexagons, as seen at right. Such hexagons are one example of a convection cell.

As the Rayleigh number is increased even further above the value where convection cells first appear, the system may undergo other bifurcations, and other more complex patterns, such as spirals, may begin to appear.

title: Convection


text:
Cryosphere

The cryosphere (from the Greek κρύος kryos, "cold", "frost" or "ice" and σφαῖρα sphaira, "globe, ball" ) is an all-encompassing term for the portions of Earth's surface where water is in solid form, including sea ice, lake ice, river ice, snow cover, glaciers, ice caps, ice sheets, and frozen ground (which includes permafrost). Thus, there is a wide overlap with the hydrosphere. The cryosphere is an integral part of the global climate system. It also has important feedbacks on the climate system. These feedbacks come from the cryosphere's influence on surface energy and moisture fluxes, clouds, the water cycle, atmospheric and oceanic circulation.

Through these feedback processes, the cryosphere plays a significant role in the global climate and in climate model response to global changes. Approximately 10% of the Earth's surface is covered by ice, but this is rapidly decreasing. Current reductions in the cryosphere (caused by climate change) are visible in ice sheet melt, glaciers decline, sea ice decline, permafrost thaw and snow cover decrease.

Overall interactions
Frozen water is found on the Earth's surface primarily as snow cover, freshwater ice in lakes and rivers, sea ice, glaciers, ice sheets, and frozen ground and permafrost (permanently frozen ground). The residence time of water in each of these cryospheric sub-systems varies widely. Snow cover and freshwater ice are essentially seasonal, and most sea ice, except for ice in the central Arctic, lasts only a few years if it is not seasonal. A given water particle in glaciers, ice sheets, or ground ice, however, may remain frozen for 10–100,000 years or longer, and deep ice in parts of East Antarctica may have an age approaching 1 million years.

Most of the world's ice volume is in Antarctica, principally in the East Antarctic Ice Sheet. In terms of areal extent, however, Northern Hemisphere winter snow and ice extent comprise the largest area, amounting to an average 23% of hemispheric surface area in January. The large areal extent and the important climatic roles of snow and ice is related to their unique physical properties. This also indicates that the ability to observe and model snow and ice-cover extent, thickness, and physical properties (radiative and thermal properties) is of particular significance for climate research.

There are several fundamental physical properties of snow and ice that modulate energy exchanges between the surface and the atmosphere. The most important properties are the surface reflectance (albedo), the ability to transfer heat (thermal diffusivity), and the ability to change state (latent heat). These physical properties, together with surface roughness, emissivity, and dielectric characteristics, have important implications for observing snow and ice from space. For example, surface roughness is often the dominant factor determining the strength of radar backscatter. Physical properties such as crystal structure, density, length, and liquid water content are important factors affecting the transfers of heat and water and the scattering of microwave energy.

The surface reflectance of incoming solar radiation is important for the surface energy balance (SEB). It is the ratio of reflected to incident solar radiation, commonly referred to as albedo. Climatologists are primarily interested in albedo integrated over the shortwave portion of the electromagnetic spectrum (~300 to 3500 nm), which coincides with the main solar energy input. Typically, albedo values for non-melting snow-covered surfaces are high (~80–90%) except in the case of forests. The higher albedos for snow and ice cause rapid shifts in surface reflectivity in autumn and spring in high latitudes, but the overall climatic significance of this increase is spatially and temporally modulated by cloud cover. (Planetary albedo is determined principally by cloud cover, and by the small amount of total solar radiation received in high latitudes during winter months.) Summer and autumn are times of high-average cloudiness over the Arctic Ocean so the albedo feedback associated with the large seasonal changes in sea-ice extent is greatly reduced. It was found that snow cover exhibited the greatest influence on Earth's radiative balance in the spring (April to May) period when incoming solar radiation was greatest over snow-covered areas.

The thermal properties of cryospheric elements also have important climatic consequences. Snow and ice have much lower thermal diffusivities than air. Thermal diffusivity is a measure of the speed at which temperature waves can penetrate a substance. Snow and ice are many orders of magnitude less efficient at diffusing heat than air. Snow cover insulates the ground surface, and sea ice insulates the underlying ocean, decoupling the surface-atmosphere interface with respect to both heat and moisture fluxes. The flux of moisture from a water surface is eliminated by even a thin skin of ice, whereas the flux of heat through thin ice continues to be substantial until it attains a thickness in excess of 30 to 40 cm. However, even a small amount of snow on top of the ice will dramatically reduce the heat flux and slow down the rate of ice growth. The insulating effect of snow also has major implications for the hydrological cycle. In non-permafrost regions, the insulating effect of snow is such that only near-surface ground freezes and deep-water drainage is uninterrupted.

While snow and ice act to insulate the surface from large energy losses in winter, they also act to retard warming in the spring and summer because of the large amount of energy required to melt ice (the latent heat of fusion, 3.34 x 105 J/kg at 0 °C). However, the strong static stability of the atmosphere over areas of extensive snow or ice tends to confine the immediate cooling effect to a relatively shallow layer, so that associated atmospheric anomalies are usually short-lived and local to regional in scale. In some areas of the world such as Eurasia, however, the cooling associated with a heavy snowpack and moist spring soils is known to play a role in modulating the summer monsoon circulation.

Climate change feedback mechanisms
There are numerous cryosphere-climate feedbacks in the global climate system. These operate over a wide range of spatial and temporal scales from local seasonal cooling of air temperatures to hemispheric-scale variations in ice sheets over time scales of thousands of years. The feedback mechanisms involved are often complex and incompletely understood. For example, Curry et al. (1995) showed that the so-called "simple" sea ice-albedo feedback involved complex interactions with lead fraction, melt ponds, ice thickness, snow cover, and sea-ice extent.

The role of snow cover in modulating the monsoon is just one example of a short-term cryosphere-climate feedback involving the land surface and the atmosphere.

Glaciers and ice sheets

Ice sheets and glaciers are flowing ice masses that rest on solid land. They are controlled by snow accumulation, surface and basal melt, calving into surrounding oceans or lakes and internal dynamics. The latter results from gravity-driven creep flow ("glacial flow") within the ice body and sliding on the underlying land, which leads to thinning and horizontal spreading. Any imbalance of this dynamic equilibrium between mass gain, loss and transport due to flow results in either growing or shrinking ice bodies.Relationships between global climate and changes in ice extent are complex. The mass balance of land-based glaciers and ice sheets is determined by the accumulation of snow, mostly in winter, and warm-season ablation due primarily to net radiation and turbulent heat fluxes to melting ice and snow from warm-air advection Where ice masses terminate in the ocean, iceberg calving is the major contributor to mass loss. In this situation, the ice margin may extend out into deep water as a floating ice shelf, such as that in the Ross Sea.

Sea ice
Sea ice covers much of the polar oceans and forms by freezing of sea water. Satellite data since the early 1970s reveal considerable seasonal, regional, and interannual variability in the sea ice covers of both hemispheres. Seasonally, sea-ice extent in the Southern Hemisphere varies by a factor of 5, from a minimum of 3–4 million km2 in February to a maximum of 17–20 million km2 in September. The seasonal variation is much less in the Northern Hemisphere where the confined nature and high latitudes of the Arctic Ocean result in a much larger perennial ice cover, and the surrounding land limits the equatorward extent of wintertime ice. Thus, the seasonal variability in Northern Hemisphere ice extent varies by only a factor of 2, from a minimum of 7–9 million km2 in September to a maximum of 14–16 million km2 in March.

The ice cover exhibits much greater regional-scale interannual variability than it does hemispherical. For instance, in the region of the Sea of Okhotsk and Japan, maximum ice extent decreased from 1.3 million km2 in 1983 to 0.85 million km2 in 1984, a decrease of 35%, before rebounding the following year to 1.2 million km2. The regional fluctuations in both hemispheres are such that for any several-year period of the satellite record some regions exhibit decreasing ice coverage while others exhibit increasing ice cover.

Snow cover
Most of the Earth's snow-covered area is located in the Northern Hemisphere, and varies seasonally from 46.5 million km2 in January to 3.8 million km2 in August.

Snow cover is an extremely important storage component in the water balance, especially seasonal snowpacks in mountainous areas of the world. Though limited in extent, seasonal snowpacks in the Earth's mountain ranges account for the major source of the runoff for stream flow and groundwater recharge over wide areas of the midlatitudes. For example, over 85% of the annual runoff from the Colorado River basin originates as snowmelt. Snowmelt runoff from the Earth's mountains fills the rivers and recharges the aquifers that over a billion people depend on for their water resources.

Furthermore, over 40% of the world's protected areas are in mountains, attesting to their value both as unique ecosystems needing protection and as recreation areas for humans.

Lake ice and river ice
Ice forms on rivers and lakes in response to seasonal cooling. The sizes of the ice bodies involved are too small to exert anything other than localized climatic effects. However, the freeze-up/break-up processes respond to large-scale and local weather factors, such that considerable interannual variability exists in the dates of appearance and disappearance of the ice. Long series of lake-ice observations can serve as a proxy climate record, and the monitoring of freeze-up and break-up trends may provide a convenient integrated and seasonally-specific index of climatic perturbations. Information on river-ice conditions is less useful as a climatic proxy because ice formation is strongly dependent on river-flow regime, which is affected by precipitation, snow melt, and watershed runoff as well as being subject to human interference that directly modifies channel flow, or that indirectly affects the runoff via land-use practices.

Lake freeze-up depends on the heat storage in the lake and therefore on its depth, the rate and temperature of any inflow, and water-air energy fluxes. Information on lake depth is often unavailable, although some indication of the depth of shallow lakes in the Arctic can be obtained from airborne radar imagery during late winter (Sellman et al. 1975) and spaceborne optical imagery during summer (Duguay and Lafleur 1997). The timing of breakup is modified by snow depth on the ice as well as by ice thickness and freshwater inflow.

Snow cover decrease
Studies in 2021 found that Northern Hemisphere snow cover has been decreasing since 1978, along with snow depth. Paleoclimate observations show that such changes are unprecedented over the last millennia in Western North America.

North American winter snow cover increased during the 20th century, largely in response to an increase in precipitation.

Because of its close relationship with hemispheric air temperature, snow cover is an important indicator of climate change.

Global warming is expected to result in major changes to the partitioning of snow and rainfall, and to the timing of snowmelt, which will have important implications for water use and management. These changes also involve potentially important decadal and longer time-scale feedbacks to the climate system through temporal and spatial changes in soil moisture and runoff to the oceans.(Walsh 1995). Freshwater fluxes from the snow cover into the marine environment may be important, as the total flux is probably of the same magnitude as desalinated ridging and rubble areas of sea ice. In addition, there is an associated pulse of precipitated pollutants which accumulate over the Arctic winter in snowfall and are released into the ocean upon ablation of the sea ice.

Related scientific disciplines
"Cryospheric sciences" is an umbrella term for the study of the cryosphere. As an interdisciplinary Earth science, many disciplines contribute to it, most notably geology, hydrology, and meteorology and climatology; in this sense, it is comparable to glaciology.

The term deglaciation describes the retreat of cryospheric features.

title: Cryosphere