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 phenomenon of the aberration of light. If we make crudely the theory of aberration, we reach a very curious result The apparent positions of the stars differ from their real positions because of the earth's motion, and as this motion is variable, these apparent positions vary. The real position we can not ascertain, but we can observe the variations of the apparent position. The observations of the aberration show us, therefore, not the earth's motion, but the variations of this motion; they can not, therefore, give us information about the absolute motion of the earth.

At least this is true in first approximation, but the case would be no longer the same if we could appreciate the thousandths of a second. Then it would be seen that the amplitude of the oscillation depends not alone on the variation of the motion, a variation which is well known, since it is the motion of our globe on its elliptic orbit, but on the mean value of this motion, so that the constant of aberration would not be quite the same for all the stars, and the differences would tell us the absolute motion of the earth in space.

This, then, would be, under another form, the ruin of the principle of relativity. We are far, it is true, from appreciating the thousandth of a second, but, after all, say some, the earth's total absolute velocity is perhaps much greater than its relative velocity with respect to the sun. If, for example, it were 300 kilometers per second in place of 30, this would suffice to make the phenomenon observable.

I believe that in reasoning thus one admits a too simple theory of aberration. Michelson has shown us, I have told you, that physical procedures are powerless to put in evidence absolute motion; I am persuaded that the same will be true of the astronomic procedures, however far precision be carried.

However that may be, the data astronomy will furnish us in this regard will some day be precious to the physicist. Meanwhile, I believe that the theorists, recalling the experience of Michelson, may anticipate a negative result, and that they would accomplish a useful work in constructing a theory of aberration which would explain this in advance.

Electrons and Spectra.—This dynamics of electrons can be approached from many sides, but among the ways leading thither is


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 one which has been somewhat neglected, and yet this is one of those which promise us the most surprises. It is movements of electrons which produce the lines of the emission spectra; this is proved by the Zeeman effect; in an incandescent body what vibrates is sensitive to the magnet, therefore electrified. This is a very important first point, but no one has gone farther. Why are the lines of the spectrum distributed in accordance with a regular law? These laws have been studied by the experimenters in their least details; they are very precise and comparatively simple. A first study of these distributions recalls the harmonics encountered in acoustics; but the difference is great. Not only are the numbers of vibrations not the successive multiples of a single number, but we do not even find anything analogous to the roots of those transcendental equations to which we are led by so many problems of mathematical physics: that of the vibrations of an elastic body of any form, that of the Hertzian oscillations in a generator of any form, the problem of Fourier for the cooling of a solid body.

The laws are simpler, but they are of wholly other nature, and to cite only one of these differences, for the harmonics of high order, the number of vibrations tends toward a finite limit, instead of increasing indefinitely.

That has not yet been accounted for, and I believe that there we have one of the most important secrets of nature. A Japanese physicist, M. Nagaoka, has recently proposed an explanation; according to him, atoms are composed of a large positive electron surrounded by a ring formed of a great number of very small negative electrons. Such is the planet Saturn with its rings. This is a very interesting attempt, but not yet wholly satisfactory; this attempt should be renewed. We will penetrate, so to speak, into the inmost recess of matter. And from the particular point of view which we to-day occupy, when we know why the vibrations of incandescent bodies differ thus from ordinary elastic vibrations, why the electrons do not behave like the matter which is familiar to us, we shall better comprehend the dynamics of electrons and it will be perhaps more easy for us to reconcile it with the principles.

Conventions Preceding Experiment.—Suppose, now, that all


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 these efforts fail, and, after all, I do not believe they will, what must be done? Will it be necessary to seek to mend the broken principles by giving what we French call a coup de pouce? That evidently is always possible, and I retract nothing of what I have said above.

Have you not written, you might say if you wished to seek a quarrel with me — have you not written that the principles, though of experimental origin, are now unassailable by experiment because they have become conventions? And now you have just told us that the most recent conquests of experiment put these principles in danger.

Well, formerly I was right and to-day I am not wrong. Formerly I was right, and what is now happening is a new proof of it. Take, for example, the calorimetric experiment of Curie on radium. Is it possible to reconcile it with the principle of the conservation of energy? This has been attempted in many ways. But there is among them one I should like you to notice; this is not the explanation which tends to-day to prevail, but it is one of those which have been proposed. It has been conjectured that radium was only an intermediary, that it only stored radiations of unknown nature which flashed through space in every direction, traversing all bodies, save radium, without being altered by this passage and without exercising any action upon them. Radium alone took from them a little of their energy and afterward gave it out to us in various forms.

What an advantageous explanation, and how convenient! First, it is unverifiable and thus irrefutable. Then again it will serve to account for any derogation whatever to Mayer's principle; it answers in advance not only the objection of Curie, but all the objections that future experimenters might accumulate. This new and unknown energy would serve for everything.

This is just what I said, and therewith we are shown that our principle is unassailable by experiment.

But then, what have we gained by this stroke? The principle is intact, but thenceforth of what use is it? It enabled us to foresee that in such or such circumstance we could count on such a total quantity of energy; it limited us; but now that this indefinite provision of new energy is placed at our disposal, we are no


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 longer limited by anything; and, as I have written in 'Science and Hypothesis,' if a principle ceases to be fecund, experiment without contradicting it directly will nevertheless have condemned it.

Future Mathematical Physics.—This, therefore, is not what would have to be done; it would be necessary to rebuild anew. If we were reduced to this necessity, we could moreover console ourselves. It would not be necessary thence to conclude that science can weave only a Penelope's web, that it can raise only ephemeral structures, which it is soon forced to demolish from top to bottom with its own hands.

As I have said, we have already passed through a like crisis. I have shown you that in the second mathematical physics, that of the principles, we find traces of the first, that of central forces; it will be just the same if we must know a third. Just so with the animal that exuviates, that breaks its too narrow carapace and makes itself a fresh one; under the new envelope one will recognize the essential traits of the organism which have persisted.

We can not foresee in what way we are about to expand; perhaps it is the kinetic theory of gases which is about to undergo development and serve as model to the others. Then the facts which first appeared to us as simple thereafter would be merely resultants of a very great number of elementary facts which only the laws of chance would make cooperate for a common end. Physical law would then assume an entirely new aspect; it would no longer be solely a differential equation, it would take the character of a statistical law.

Perhaps, too, we shall have to construct an entirely new mechanics that we only succeed in catching a glimpse of, where, inertia increasing with the velocity, the velocity of light would become an impassable limit. The ordinary mechanics, more simple, would remain a first approximation, since it would be true for velocities not too great, so that the old dynamics would still be found under the new. We should not have to regret having believed in the principles, and even, since velocities too great for the old formulas would always be only exceptional, the surest way in practise would be still to act as if we continued to


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 believe in them. They are so useful, it would be necessary to keep a place for them. To determine to exclude them altogether would be to deprive oneself of a precious weapon. I hasten to say in conclusion that we are not yet there, and as yet nothing proves that the principles will not come forth from out the fray victorious and intact.


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 read with much interest Messrs. Michelson and Morley's wonderfully delicate experiment attempting to decide the important question as to how far the ether is carried along by the earth. Their result seems opposed to other experiments showing that the ether in the air can be carried along only to an inappreciable extent. I would suggest that almost the only hypothesis that can reconcile this opposition is that the length of material bodies changes, according as they are moving through the ether or across it, by an amount depending on the square of the ratio of their velocity to that of light. We know that electric forces are affected by the motion of the electrified bodies relative to the ether, and it seems a not improbable supposition that the molecular forces are affected by the motion, and that the size of a body alters consequently. It would be very important if secular experiments on electrical attractions between permanently electrified bodies, such as in a very delicate quadrant electrometer, were instituted in some of the equatorial parts of the earth to observe whether there is any diurnal and annual variation of attraction, — diurnal due to the rotation of the earth being added and subtracted from its orbital velocity; and annual similarly for its orbital velocity and the motion of the solar system.

Dublin, May 2.

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CHAPTER XIII

THE GALAPAGOS ISLANDS

 * Position and Physical Features—Absence of Indigenous Mammalia and Amphibia—Reptiles—Birds—Insects and Land-Shells—The Keeling Islands as Illustrating the Manner in which Oceanic Islands are Peopled—Flora of the Galapagos—Origin of the Flora of the Galapagos—Concluding Remarks.

The Galapagos differ in many important respects from the islands we have examined in our last chapter, and the differences are such as to have affected the whole character of their animal inhabitants. Like the Azores, they are volcanic, but they are much more extensive, the islands being both larger and more numerous; while volcanic action has been so recent that a large portion of their surface consists of barren lava-fields. They are considerably less distant from a continent than either the Azores or Bermuda, being about 600 miles from the west coast of South America and a little more than 700 from Veragua, with the small Cocos Islands intervening; and they are situated on the equator instead of being in the north temperate zone. They stand upon a deeply submerged bank, the 1,000 fathom line encircling all the more important islands at a few miles distance, whence there appears to be a comparatively steep descent all round to the average depth of that portion of the Pacific, between 2,000 and 3,000 fathoms.

The whole group occupies a space of about 300 by 200 miles. It consists of five large and twelve small islands; the largest (Albemarle Island) being about eighty miles long and of very irregular shape, while the four next in importance—Chatham, Indefatigable, James, and Narborough Islands, are each about twenty-five or thirty miles long, and of a rounded or elongate form. The whole are entirely volcanic, and in the western islands there are numerous active volcanoes. Unlike the other groups of islands we have been considering, these are situated in a comparatively calm sea, where storms are of rare occurrence and even strong winds almost unknown. They are traversed by ocean currents which are strong and constant, flowing towards the north-west from the coast of Peru; and these physical conditions have had a powerful influence on the animal and vegetable forms by which the islands are now inhabited. The Galapagos have also, during three centuries, been frequently visited by Europeans, and were long a favourite resort of buccaneers and traders, who found an ample supply of food in the large tortoises which abound there; and to these visits we may perhaps trace the introduction of some animals whose presence it is otherwise difficult to account for. The vegetation is generally scanty, but still amply sufficient for the support of a considerable amount of animal life, as shown by the cattle, horses, asses, goats, pigs, dogs, and cats, which now run wild in some of the islands.

Absence of Indigenous Mammalia and Amphibia.—As in all other oceanic islands, we find here no truly indigenous mammalia, for though there is a mouse of the American genus Hesperomys, which differs somewhat from any known species, we can hardly consider this to be indigenous; first, because these creatures have been little studied in South America, and there may yet be many undescribed species, and in the second place because even had it been introduced by some European or native vessel, there is ample time in two or three hundred years for the very different conditions to have established a marked diversity in the characters of the species. This is the more probable because there is also a true rat of the Old World genus Mus, which is said to differ slightly from any known species; and as this genus is not a native of the American continents we are sure that it must have been recently introduced into the Galapagos. There can be little doubt therefore that the islands are completely destitute of truly indigenous mammalia; and frogs and toads, the only tropical representatives of the Amphibia, are equally unknown.

Reptiles.—Reptiles, however, which at first sight appear as unsuited as mammals to pass over a wide expanse of ocean, abound in the Galapagos, though the species are not very numerous. They consist of land-tortoises, lizards and snakes. The tortoises consist of two peculiar species, Testudo microphyes, found in most of the islands, and T. abingdonii recently discovered on Abingdon Island, as well as one extinct species, T. ephippium, found on Indefatigable Island. These are all of very large size, like the gigantic tortoises of the Mascarene Islands, from which, however, they differ in structural characters; and Dr. Günther believes that they have been originally derived from the American continent. Considering the well known tenacity of life of these animals, and the large number of allied forms which have aquatic or sub-aquatic habits, it is not a very extravagant supposition that some ancestral form, carried out to sea by a flood, was once or twice safely drifted as far as the Galapagos, and thus originated the races which now inhabit them.

The lizards are five in number; a peculiar species of gecko, Phyllodactylus galapagensis, and four species of the American family Iguanidæ. Two of these are distinct species of the genus Tropidurus, the other two being large, and so very distinct as to be classed in peculiar genera. One of these is aquatic and found in all the islands, swimming in the sea at some distance from the shore and feeding on seaweed; the other is terrestrial, and is confined to the four central islands. These last were originally described as Amblyrhynchus cristatus by Mr. Bell, and A. subcristatus by Gray; they were afterwards placed in two other genera Trachycephalus and Oreocephalus (see Brit. Mus. Catalogue of Lizards), while in a recent paper by Dr. Steindachner, the marine species is again classed as Amblyrhynchus, while the terrestrial form is placed in another genus Conolophus, both genera being peculiar to the Galapagos.

How these lizards reached the islands we cannot tell. The fact that they all belong to American genera or families indicates their derivation from that continent, while their being all distinct species is a proof that their arrival took place at a remote epoch, under conditions perhaps somewhat different from any which now prevail. It is certain that animals of this order have some means of crossing the sea not possessed by any other land vertebrates, since they are found in a considerable number of islands which possess no mammals nor any other land reptiles; but what those means are has not yet been positively ascertained.

It is unusual for oceanic islands to possess snakes, and it is therefore somewhat of an anomaly that two species are found in the Galapagos. Both are closely allied to South American forms, and one is hardly different from a Chilian snake, so that they indicate a more recent origin than in the case of the lizards. Snakes it is known can survive a long time at sea, since a living boa-constrictor once reached the island of St. Vincent from the coast of South America, a distance of two hundred miles by the shortest route. Snakes often frequent trees, and might thus be conveyed long distances if carried out to sea on a tree uprooted by a flood such as often occurs in tropical climates and especially during earthquakes. To some such accident we may perhaps attribute the presence of these creatures in the Galapagos, and that it is a very rare one is indicated by the fact that only two species have as yet succeeded in obtaining a footing there.

Birds.—We now come to the birds, whose presence here may not seem so remarkable, but which yet present features of interest not exceeded by any other group. About seventy species of birds have now been obtained on these islands, and of these forty-one are peculiar to them. But all the species found elsewhere, except one, belong to the aquatic tribes or the waders which are pre-eminently wanderers, yet even of these eight are peculiar. The true land-birds are forty-two in number, and all but one are entirely confined to the Galapagos; while three-fourths of them present such peculiarities that they are classed in distinct genera. All are allied to birds inhabiting tropical America, some very closely; while one—the common American rice-bird which ranges over the whole northern and part of the southern continents—is the only land-bird identical with those of the mainland. The following is a list of these land-birds taken from Mr. Salvin's memoir in the Transactions of the Zoological Society for the year 1876, to which are added nine species collected in 1888 and described by Mr. Ridgway in the Proceedings of the U.S. National Museum (XII. p. 101) and some additional species obtained in 1889.

We have here every gradation of difference from perfect identity with the continental species to genera so distinct that it is difficult to determine with what forms they are most nearly allied; and it is interesting to note that this diversity bears a distinct relation to the probabilities of, and facilities for, migration to the islands. The excessively abundant rice-bird, which breeds in Canada and swarms over the whole United States, migrating to the West Indies and South America, visiting the distant Bermudas almost every year, and extending its range as far as Paraguay, is the only species of land-bird which remains completely unchanged in the Galapagos; and we may therefore conclude that some stragglers of the migrating host reach the islands sufficiently often to keep up the purity of the breed. Next, we have the almost cosmopolite short-eared owl (Asio brachyotus), which ranges from China to Ireland, and from Greenland to the Straits of Magellan, and of this the Galapagos bird is probably only one of the numerous varieties. The little wood warbler (Dendrœca aureola) is closely allied to a species which ranges over the whole of North America and as far south as New Grenada. It has also been occasionally met with in Bermuda, an indication that it has considerable powers of flight and endurance. The more distinct species—as the tyrant fly-catchers (Pyrocephalus and Myiarchus), the ground-dove (Zenaida), and the buzzard (Buteo), are all allied to non-migratory species peculiar to tropical America, and of a more restricted range; while the distinct genera are allied to South American groups of thrushes, finches, and sugar-birds which have usually restricted ranges, and whose habits are such as not to render them likely to be carried out to sea. The remote ancestral forms of these birds which, owing to some exceptional causes, reached the Galapagos, have thus remained uninfluenced by later migrations, and have, in consequence, been developed into a variety of distinct types adapted to the peculiar conditions of existence under which they have been placed. Sometimes the different species thus formed are confined to one or two of the islands only, as the three species of Certhidea, which are divided between the islands but do not appear ever to occur together. Nesomimus parvulus is confined to Albemarle Island, and N. trifasciatus to Charles Island; Cactornis pallida to Indefatigable Island, C. brevirostris to Chatham Island, and C. abingdoni to Abingdon Island.

Now all these phenomena are strictly consistent with the theory of the peopling of the islands by accidental migrations, if we only allow them to have existed for a sufficiently long period; and the fact that volcanic action has ceased on many of the islands, as well as their great extent, would certainly indicate a considerable antiquity.

The great difference presented by the birds of these islands as compared with those of the equally remote Azores and Bermudas, is sufficiently explained by the difference of climatal conditions. At the Galapagos there are none of those periodic storms, gales, and hurricanes which prevail in the North Atlantic, and which every year carry some straggling birds of Europe or North America to the former islands; while, at the same time, the majority of the tropical American birds are nonmigratory, and thus afford none of the opportunities presented by the countless hosts of migrants which pass annually northward and southward along the European, and especially along the North American coasts. It is strictly in accordance with these different conditions that we find in one case an almost perfect identity with, and in the other an almost equally complete diversity from, the continental species of birds.

Insects and Land-shells.—The other groups of land-animals add little of importance to the facts already referred to. The insects are very scanty; the most plentiful group, the beetles, only furnishing about forty species belonging to thirty-two genera and nineteen families. The species are almost all peculiar, as are some of the genera. They are mostly small and obscure insects, allied either to American or to world-wide groups. The Carabidæ and the Heteromera are the most abundant groups, the former furnishing six and the latter nine species.

The land-shells are not abundant—about twenty in all, most of them peculiar species, but not otherwise remarkable. The observation of Captain Collnet, quoted by Mr. Darwin in his Journal, that drift-wood, bamboos, canes, and the nuts of a palm, are often washed on the south-eastern shores of the islands, furnishes an excellent clue to the manner in which many of the insects and land-shells may have reached the Galapagos. Whirlwinds also have been known to carry quantities of leaves and other vegetable débris to great heights in the air, and these might be then carried away by strong upper currents and dropped at great distances, and with them small insects and mollusca, or their eggs. We must also remember that volcanic islands are subject to subsidence as well as elevation; and it is quite possible that during the long period the Galapagos have existed some islands may have intervened between them and the coast, and have served as stepping-stones by which the passage to them of various organisms would be greatly facilitated. Sunken banks, the relics of such islands, are known to exist in many parts of the ocean, and countless others, no doubt, remain undiscovered.

The Keeling Islands as Illustrating the Manner in which Oceanic Islands are Peopled.—That such causes as have been here adduced are those by which oceanic islands have been peopled, is further shown by the condition of equally remote islands which we know are of comparatively recent origin. Such are the Keeling or Cocos Islands in the Indian Ocean, situated about the same distance from Sumatra as the Galapagos from South America, but mere coral reefs, supporting abundance of cocoa-nut palms as their chief vegetation. These islands were visited by Mr. Darwin, and their natural history carefully examined. The only mammals are rats, brought by a wrecked vessel and said by Mr. Waterhouse to be common English rats, "but smaller and more brightly coloured;" so that we have here an illustration of how soon a difference of race is established under a constant and uniform difference of conditions. There are no true land-birds, but there are snipes and rails, both apparently common Malayan species. Reptiles are represented by one small lizard, but no account of this is given in the Zoology of the Voyage of the Beagle, and we may therefore conclude that it was an introduced species. Of insects, careful collecting only produced thirteen species belonging to eight distinct orders. The only beetle was a small Elater, the Orthoptera were a Gryllus and a Blatta; and there were two flies, two ants, and two small moths, one a Diopæa which swarms everywhere in the eastern tropics in grassy places. All these insects were no doubt brought either by winds, by floating timber (which reaches the islands abundantly), or by clinging to the feathers of aquatic or wading birds; and we only require more time to introduce a greater variety of species, and a better soil and more varied vegetation, to enable them to live and multiply, in order to give these islands a fauna and flora equal to that of the Bermudas. Of wild plants there were only twenty species, belonging to nineteen genera and to no less than sixteen natural families, while all were common tropical shore plants. These islands are thus evidently stocked by waifs and strays brought by the winds and waves; but their scanty vegetation is mainly due to unfavourable conditions—the barren coral rock and sand, of which they are wholly composed, together with exposure to sea-air, being suitable to a very limited number of species which soon monopolise the surface. With more variety of soil and aspect a greater variety of plants would establish themselves, and these would favour the preservation and increase of more insects, birds, and other animals, as we find to be the case in many small and remote islands.

Flora of the Galapagos.—The plants of these islands are so much more numerous than the known animals, even including the insects, they have been so carefully studied by eminent botanists, and their relations throw so much light on the past history of the group, that no apology is needed for giving a brief outline of the peculiarities and affinities of the flora. The statements we shall make on this subject will be taken from the Memoir of Sir Joseph Hooker in the Linnæan Transactions for 1851, founded on Mr. Darwin's collections, and a later paper by N. J. Andersson in the Linnæa of 1861, embodying more recent discoveries.

The total number of flowering plants known at the latter date was 332, of which 174 were peculiar to the islands, while 158 were common to other countries. Of these latter about twenty have been introduced by man, while the remainder are all natives of some part of America, though about a third part are species of wide range extending into both hemispheres. Of those confined to America, forty-two are found in both the northern and southern continents, twenty-one are confined to South America, while twenty are found only in North America, the West Indies, or Mexico. This equality of North American and South American species in the Galapagos is a fact of great significance in connection with the observation of Sir Joseph Hooker that the peculiar species are allied to the plants of temperate America or to those of the high Andes, while the non-peculiar species are mostly such as inhabit the hotter regions of the tropics near the level of the sea. He also observes that the seeds of this latter class of Galapagos plants often have special means of transport, or belong to groups whose seeds are known to stand long voyages and to possess great vitality. Mr. Bentham also, in his elaborate account of the Compositæ, remarks on the decided Central American or Mexican affinities of the Galapagos species, so that we may consider this to be a thoroughly well-established fact.

The most prevalent families of plants in the Galapagos are the Compositæ (40 sp.), Gramineæ (32 sp.), Leguminosæ (30 sp.), and Euphorbiaceæ (29 sp.). Of the Compositæ most of the species, except such as are common weeds or shore plants, are peculiar, but there are only two peculiar genera, allied to Mexican forms and not very distinct; while the genus Lipochæta, represented here by a single species, is only found elsewhere in the Sandwich Islands though it has American affinities.

Origin of the Galapagos Flora.—These facts are explained by the past history of the American continent, its separation at various epochs by arms of the sea uniting the two oceans across what is now Central America (the last separation being of recent date, as shown by the considerable number of identical species of fishes on both sides of the isthmus), and the influence of the glacial epoch in driving the temperate American flora southward along the mountain plateaus. At the time when the two oceans were united a portion of the Gulf Stream may have been diverted into the Pacific, giving rise to a current, some part of which would almost certainly have reached the Galapagos, and this may have helped to bring about that singular assemblage of West Indian and Mexican plants now found there. And as we now believe that the duration of the last glacial epoch in its successive phases was much longer than the time which has elapsed since it finally passed away, while throughout the Miocene epoch the snow-line would often be lowered during periods of high excentricity, we are enabled to comprehend the nature of the causes which may have led to the islands being stocked with those north tropical or mountain types which are so characteristic a feature of that portion of the Galapagos flora which consists of peculiar species.

On the whole, the flora agrees with the fauna in indicating a moderately remote origin, great isolation, and changes of conditions affording facilities for the introduction of organisms from various parts of the American coast, and even from the West Indian Islands and Gulf of Mexico. As in the case of the birds, the several islands differ considerably in their native plants, many species being limited to one or two islands only, while others extend to several. This is, of course, what might be expected on any theory of their origin; because, even if the whole of the islands had once been united and afterwards separated, long continued isolation would often lead to the differentiation of species, while the varied conditions to be found upon islands differing in size and altitude as well as in luxuriance of vegetation, would often lead to the extinction of a species on one island and its preservation on another. If the several islands had been equally well explored, it might be interesting to see whether, as in the case of the Azores, the number of species diminished in those more remote from the coast; but unfortunately our knowledge of the productions of the various islands of the group is exceedingly unequal, and, except in those cases in which representative species inhabit distinct islands, we have no certainty on the subject. All the more interesting problems in geographical distribution, however, arise from the relation of the fauna and flora of the group as a whole to those of the surrounding continents, and we shall therefore for the most part confine ourselves to this aspect of the question in our discussion of the phenomena presented by oceanic or continental islands.

Concluding Remarks.—The Galapagos offer an instructive contrast with the Azores, showing how a difference of conditions that might be thought unimportant may yet produce very striking results in the forms of life. Although the Galapagos are much nearer a continent than the Azores, the number of species of plants common to the continent is much less in the former case than in the latter, and this is still more prominent a characteristic of the insect and the bird faunas. This difference has been shown to depend, almost entirely, on the one archipelago being situated in a stormy, the other in a calm portion of the ocean; and it demonstrates the preponderating importance of the atmosphere as an agent in the dispersal of birds, insects, and plants. Yet ocean-currents and surface-drifts are undoubtedly efficient carriers of plants, and, with plants, of insects and shells, especially in the tropics; and it is probably to this agency that we may impute the recent introduction of a number of common Peruvian and Chilian littoral species, and also of several West Indian types at a more remote period when the Isthmus of Panama was submerged.

In the case of these islands we see the importance of taking account of past conditions of sea and land and past changes of climate, in order to explain the relations of the peculiar or endemic species of their fauna and flora; and we may even see an indication of the effects of climatal changes in the northern hemisphere, in the north temperate or alpine affinities of many of the plants, and even of some of the birds. The relation between the migratory habits of the birds and the amount of difference from continental types is strikingly accordant with the fact that it is almost exclusively migratory birds that annually reach the Azores and Bermuda; while the corresponding fact that the seeds of those plants, which are common to the Galapagos and the adjacent continent, have all—as Sir Joseph Hooker states—some special means of dispersal, is equally intelligible. The reason why the Galapagos possess four times as many peculiar species of plants as the Azores is clearly a result of the less constant introduction of seeds, owing to the absence of storms; the greater antiquity of the group, allowing more time for specific change; and the influence of cold epochs and of alterations of sea and land, in bringing somewhat different sets of plants at different times within the influence of such modified winds and currents as might convey them to the islands.

On the whole, then, we have no difficulty in explaining the probable origin of the flora and fauna of the Galapagos, by means of the illustrative facts and general principles already adduced.

112 Gigantic Land Tortoises Living and Extinct in the Collection of the British Museum. By A. C. L. G. Günther, F.R.S. 1877.

113 The following list of the beetles yet known from the Galapagos shows their scanty proportions and accidental character; the forty species belonging to thirty-three genera and eighteen families. It is taken from Mr. Waterhouse's enumeration in the Proceedings of the Zoological Society for 1877 (p. 81), with a few additions collected by the U. S. Fish Commission Steamer Albatross, and published by the U. S. National Museum in 1889.

114 Mr. H. O. Forbes, who visited these islands in 1878, increased the number of wild plants to thirty-six, and these belonged to twenty-six natural orders.

115 Juan Fernandez is a good example of a small island which, with time and favourable conditions, has acquired a tolerably rich and highly peculiar flora and fauna. It is situated in 34° S. Lat., 400 miles from the coast of Chile, and so far as facilities for the transport of living organisms are concerned is by no means in a favourable position, for the ocean-currents come from the south-west in a direction where there is no land but the Antarctic continent, and the prevalent winds are also westerly. No doubt, however, there are occasional storms, and there may have been intermediate islands, but its chief advantages are its antiquity, its varied surface, and its favourable soil and climate, offering many chances for the preservation and increase of whatever plants and animals have chanced to reach it. The island consists of basalt, greenstone, and other ancient rocks, and though only about twelve miles long its mountains are three thousand feet high. Enjoying a moist and temperate climate it is especially adapted to the growth of ferns, which are very abundant; and as the spores of these plants are as fine as dust, and very easily carried for enormous distances by winds, it is not surprising that there are nearly fifty species on the island, while the remote period when it first received its vegetation may be indicated by the fact that nearly half the species are quite peculiar; while of 102 species of flowering plants seventy are peculiar, and there are ten peculiar genera. The same general character pervades the fauna. For so small an island it is rich, containing four true land-birds, about fifty species of insects, and twenty of land-shells. Almost all these belong to South American genera, and a large proportion are South American species; but several of the insects, half the birds, and the whole of the land-shells are peculiar. This seems to indicate that the means of transmission were formerly greater than they are now, and that in the case of land-shells none have been introduced for so long a period that all have become modified into distinct forms, or have been preserved on the island while they have become extinct on the continent. For a detailed examination of the causes which have led to the modification of the humming birds of Juan Fernandez see the chapter on Humming Birds in the author's Natural Selection and Tropical Nature, p. 324; while a general account of the fauna of the island is given in his Geographical Distribution of Animals, Vol. II. p. 49.

116 No additions appear to have been made to this flora down to 1885, when Mr. Hemsley published his Report on the Present State of our Knowledge of Insular Floras.

117 Journal of the Linnean Society, Vol. XIII., "Botany," p. 556.

118 Geographical Distribution of Animals, Vol. II. p. 81.

title: Island Life/XIII


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 Rh improve; everything is elegant, clean, and neat; the streets are widened where they were narrow; even Snow Hill is become almost level, and is a very handsome street; and the narrow part of the Strand near St. Clement s is widened and become very elegant.

My wife continues poorly, but fancies she is better in health here than by the seaside. We both sincerely pray for the health of Miss Poole, and for all our friends in Sussex, and remain, dear sir, your sincere and devoted servants,

W. and C..

29.

From to.

,

2nd January 1804.

,—Mr. Blake s opinion that the drawing sent from Norfolk may be advantageously engraved for the ensuing volume of Cowper's Life, as an agreeable perspective of the situation, seems very just, whilst the Monument itself may be represented on a larger scale in a vignette, and for the materials on this subject, he


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 134 will be at no loss. I sincerely wish with you that the trial was over, that our poor friend's peace of mind might be restored; although I have no doubt from what I have heard of the soldier's character and the merits of the case, that the bill will at least be thrown out by the Court as groundless and vexatious. Blake's irritability, as well as the association and arrangement of his ideas, do not seem likely to be soothed or more advantageously disposed by any power inferior to That by which man is originally endowed with his faculties. I wish all our defects were fewer; certainly my own among the rest. But if we really are desirous this should come to pass, we are told to whom and by what means we should apply.

I wonder, my good friend, as you admired the genius of Romney so much, that you do not remember the whole catalogue of his chalk cartoons; as I think it was your opinion, in common with other sufficient judges, that they were the noblest of his studies. Besides, they were but few in number. The following were the subjects: "A Lapland Witch raising a Storm"; "Charity and her Children"; "Pliny and his Mother flying from

Dereham, containing the Grave of William Cowper, Esq." Francis Stone, del.; W. Blake, sculpt. (2) Plate facing p. 416, "A Sketch of the Monument Erected in the Church of East Dereham in Norfolk, in Memory of William Cowper, Esqre." Etch'd by W. Blake from the original Model by John Flaxman.


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Island Life/XIV

CHAPTER XIV

ST. HELENA

 * Position and Physical Features of St. Helena—Change Effected by European Occupation—The Insects of St. Helena—Coleoptera—Peculiarities and Origin of the Coleoptera of St. Helena—Land-shells of St. Helena—Absence of Fresh-water Organisms—Native Vegetation of St. Helena—The Relations of the St. Helena Compositæ—Concluding Remarks on St. Helena.

In order to illustrate as completely as possible the peculiar phenomena of oceanic islands, we will next examine the organic productions of St. Helena and of the Sandwich Islands, since these combine in a higher degree than any other spots upon the globe, extreme isolation from all more extensive lands, with a tolerably rich fauna and flora whose peculiarities are of surpassing interest. Both, too, have received considerable attention from naturalists; and though much still remains to be done in the latter group, our knowledge is sufficient to enable us to arrive at many interesting results.

Position and Physical Features of St. Helena.—This island is situated nearly in the middle of the South Atlantic Ocean, being more than 1,100 miles from the coast of Africa, and 1,800 from South America. It is about ten miles long by eight wide, and is wholly volcanic, consisting of ancient basalts, lavas, and other volcanic products. It is very mountainous and rugged, bounded for the most part by enormous precipices, and rising to a height of 2,700 feet above the sea-level. An ancient crater, about four miles across, is open on the south side, and its northern rim forms the highest and central ridge of the island. Many other hills and peaks, however, are more than two thousand feet high, and a considerable portion of the surface consists of a rugged plateau, having an elevation of about fifteen hundred to two thousand feet. Everything indicates that St. Helena is an isolated volcanic mass built up from the depths of the ocean. Mr. Wollaston remarks: "There are the strongest reasons for believing that the area of St. Helena was never very much larger than it is at present—the comparatively shallow sea-soundings within about a mile and a half from the shore revealing an abruptly defined ledge, beyond which no bottom is reached at a depth of 250 fathoms; so that the original basaltic mass, which was gradually piled up by means of successive eruptions from beneath the ocean, would appear to have its limit definitely marked out by this suddenly-terminating submarine cliff—the space between it and the existing coast-line being reasonably referred to that slow process of disintegration by which the island has been reduced, through the eroding action of the elements, to its present dimensions." If we add to this that between the island and the coast of Africa, in a south-easterly direction, is a profound oceanic gulf known to reach a depth of 2,860 fathoms, or 17,160 feet, while an equally deep, or perhaps deeper, ocean, extends to the west and south-west, we shall be satisfied that St. Helena is a true oceanic island, and that it owes none of its peculiarities to a former union with any continent or other distant land.

Change Effected by European Occupation.—When first discovered, in the year 1501, St. Helena was densely covered with a luxuriant forest vegetation, the trees overhanging the seaward precipices and covering every part of the surface with an evergreen mantle. This indigenous vegetation has been almost wholly destroyed; and although an immense number of foreign plants have been introduced, and have more or less completely established themselves, yet the general aspect of the island is now so barren and forbidding that some persons find it difficult to believe that it was once all green and fertile. The cause of the change is, however, very easily explained. The rich soil formed by decomposed volcanic rock and vegetable deposits could only be retained on the steep slopes so long as it was protected by the vegetation to which it in great part owed its origin. When this was destroyed, the heavy tropical rains soon washed away the soil, and has left a vast expanse of bare rock or sterile clay. This irreparable destruction was caused in the first place by goats, which were introduced by the Portuguese in 1513, and increased so rapidly that in 1588, they existed in thousands. These animals are the greatest of all foes to trees, because they eat off the young seedlings, and thus prevent the natural restoration of the forest. They were, however, aided by the reckless waste of man. The East India Company took possession of the island in 1651, and about the year 1700 it began to be seen that the forests were fast diminishing, and required some protection. Two of the native trees, redwood and ebony, were good for tanning, and to save trouble the bark was wastefully stripped from the trunks only, the remainder being left to rot; while in 1709 a large quantity of the rapidly disappearing ebony was used to burn lime for building fortifications! By the MSS. records quoted in Mr. Melliss' interesting volume on St. Helena, it is evident that the evil consequences of allowing the trees to be destroyed were clearly foreseen, as the following passages show: "We find the place called the Great Wood in a flourishing condition, full of young trees, where the hoggs (of which there is a great abundance) do not come to root them up. But the Great Wood is miserably lessened and destroyed within our memories, and is not near the circuit and length it was. But we believe it does not contain now less than fifteen hundred acres of fine woodland and good ground, but no springs of water but what is salt or brackish, which we take to be the reason that that part was not inhabited when the people first chose out their settlements and made plantations; but if wells could be sunk, which the governor says he will attempt when we have more hands, we should then think it the most pleasant and healthiest part of the island. But as to healthiness, we don't think it will hold so if the wood that keeps the land warm were destroyed, for then the rains, which are violent here, would carry away the upper soil, and it being a clay marl underneath would produce but little; as it is, we think in case it were enclosed it might be greatly improved" ... "When once this wood is gone the island will soon be ruined" ... "We viewed the wood's end which joins the Honourable Company's plantation called the Hutts, but the wood is so destroyed that the beginning of the Great Wood is now a whole mile beyond that place, and all the soil between being washed away, that distance is now entirely barren." (MSS. records, 1716.) In 1709 the governor reported to the Court of Directors of the East India Company that the timber was rapidly disappearing, and that the goats should be destroyed for the preservation of the ebony wood, and because the island was suffering from droughts. The reply was, "The goats are not to be destroyed, being more valuable than ebony." Thus, through the gross ignorance of those in power, the last opportunity of preserving the peculiar vegetation of St. Helena, and preventing the island from becoming the comparatively rocky desert it now is, was allowed to pass away. Even in a mere pecuniary point of view the error was a fatal one, for in the next century (in 1810) another governor reports the total destruction of the great forests by the goats, and that in consequence the cost of importing fuel for government use was 2,729l. 7s. 8d. for a single year! About this time large numbers of European, American, Australian, and South African plants were imported, and many of these ran wild and increased so rapidly as to drive out and exterminate much of the relics of the native flora; so that now English broom gorse and brambles, willows and poplars, and some common American, Cape, and Australian weeds, alone meet the eye of the ordinary visitor. These, in Sir Joseph Hooker's opinion, render it absolutely impossible to restore the native flora, which only lingers in a few of the loftiest ridges and most inaccessible precipices, and is rarely seen except by some exploring naturalist.

This almost total extirpation of a luxuriant and highly peculiar vegetation must inevitably have caused the destruction of a considerable portion of the lower animals which once existed on the island, and it is rather singular that so much as has actually been discovered should be left to show us the nature of the aboriginal fauna. Many naturalists have made small collections during short visits, but we owe our present complete knowledge of the two most interesting groups of animals, the insects, and the land-shells, mainly to the late Mr. T. Vernon Wollaston, who, after having thoroughly explored Madeira and the Canaries, undertook a voyage to St. Helena for the express purpose of studying its terrestrial fauna, and resided for six months (1875-76) in a high central position, whence the loftiest peaks could be explored. The results of his labours are contained in two volumes, which, like all that he wrote, are models of accuracy and research, and it is to these volumes that we are indebted for the interesting and suggestive facts which we here lay before our readers.

Insects—Coleoptera.—The total number of species of beetles hitherto observed at St. Helena is 203, but of these no less than seventy-four are common and wide-spread insects, which have certainly, in Mr. Wollaston's opinion, been introduced by human agency. There remain 129 which are believed to be truly aborigines, and of these all but one are found nowhere else on the globe. But in addition to this large amount of specific peculiarity (perhaps unequalled anywhere else in the world) the beetles of this island are equally remarkable for their generic isolation, and for the altogether exceptional proportion in which the great divisions of the order are represented. The species belong to thirty-nine genera, of which no less than twenty-five are peculiar to the island; and many of these are such isolated forms that it is impossible to find their allies in any particular country. Still more remarkable is the fact, that more than two-thirds of the whole number of indigenous species are Rhyncophora or weevils, while more than two-fifths (fifty-four species) belong to one family, the Cossonidæ. Now although the Rhyncophora are an immensely numerous group and always form a large portion of the insect population, they nowhere else approach such a proportion as this. For example, in Madeira they form one-sixth of the whole of the indigenous Coleoptera, in the Azores less than one-tenth, and in Britain one-seventh. Even more interesting is the fact that the twenty genera to which these insects belong are every one of them peculiar to the island, and in many cases have no near allies elsewhere, so that we cannot but look on this group of beetles as forming the most characteristic portion of the ancient insect fauna. Now, as the great majority of these are wood borers, and all are closely attached to vegetation and often to particular species of plants, we might, as Mr. Wollaston well observes, deduce the former luxuriant vegetation of the island from the great preponderance of this group, even had we not positive evidence that it was at no distant epoch densely forest-clad. We will now proceed briefly to indicate the numbers and peculiarities of each of the families of beetles which enter into the St. Helena fauna, taking them, not in systematic order, but according to their importance in the island.

—This great division includes the weevils and allied groups, and, as above stated, exceeds in number of species all the other beetles of the island. Four families are represented; the Cossonidæ, with fifteen peculiar genera comprising fifty-four species, and one minute insect (Stenoscelis hylastoides) forming a peculiar genus, but which has been found also at the Cape of Good Hope. It is therefore impossible to say of which country it is really a native, or whether it is indigenous to both, and dates back to the remote period when St. Helena received its early emigrants. All the Cossonidæ are found in the highest and wildest parts of the island where the native vegetation still lingers, and many of them are only found in the decaying stems of tree-ferns, box-wood, arborescent Compositæ, and other indigenous plants. They are all pre-eminently peculiar and isolated, having no direct affinity to species found in any other country. The next family, the Tanyrhynchidæ, has one peculiar genus in St. Helena, with ten species. This genus (Nesiotes) is remotely allied to European, Australian, and Madeiran insects of the same family: the habits of the species are similar to those of the Cossonidæ. The Trachyphlœidæ are represented by a single species belonging to a peculiar genus not very remote from a European form. The Anthribidæ again are highly peculiar. There are twenty-six species belonging to three genera, all endemic, and so extremely peculiar that they form two new subfamilies. One of the genera, Acarodes, is said to be allied to a Madeiran genus.

—These are the terrestrial carnivorous beetles, very abundant in all parts of the world, especially in the temperate regions of the northern hemisphere. In St. Helena there are fourteen species belonging to three genera, one of which is peculiar. This is the Haplothorax burchellii, the largest beetle on the island, and now very rare. It resembles a large black Carabus. There is also a peculiar Calosoma, very distinct, though resembling in some respects certain African species. The rest of the Geodephaga, twelve in number, belong to the wide-spread genus Bembidium, but they are altogether peculiar and isolated, except one, which is of European type, and alone has wings, all the rest being wingless.

—This group is represented by three peculiar genera containing four species, with two species belonging to European genera. They belong to the families Opatridæ, Mordellidæ, and Anthicidæ.

—Of this group there are six peculiar species belonging to four European genera—Homalota, Philonthus, Xantholinus, and Oxytelus.

—The families Elateridæ and Anobiidæ are each represented by a peculiar species of a European genus.

—There are only three species of this tribe, belonging to the European genus Longitarsus.

—Here are three species belonging to two genera. One is a peculiar species of Trox, allied to South African forms; the other two belong to the peculiar genus Melissius, which Mr. Wollaston considers to be remotely allied to Australian insects.

—Here we have the fine lady-bird Chilomenus lunata, also found in Africa, but apparently indigenous in St. Helena; and a peculiar species of Euxestes, a genus only found elsewhere in Madeira.

—These, the minutest of beetles, are represented by one species of the European and Madeiran genus Ptinella.

—One indigenous species of Cryptophaga inhabits St. Helena, and this is said to be very closely allied to a Cape species.

Peculiarities and Origin of the Coleoptera of St. Helena.—We see that the great mass of the indigenous species are not only peculiar to the island, but so isolated in their characters as to show no close affinity with any existing insects; while a small number (about one-third of the whole) have some relations, though often very remote, with species now inhabiting Europe, Madeira, or South Africa. These facts clearly point to the very great antiquity of the insect fauna of St. Helena, which has allowed time for the modification of the originally introduced species, and their special adaptation to the conditions prevailing in this remote island. This antiquity is also shown by the remarkable specific modification of a few types. Thus the whole of the Cossonidæ may be referred to three types, one species only (Hexacoptus ferrugineus) being allied to the European Cossonidæ though forming a distinct genus; a group of three genera and seven species remotely allied to the Stenoscelis hylastoides, which occurs also at the Cape; while a group of twelve genera with forty-six species have their only (remote) allies in a few insects widely scattered in South Africa, New Zealand, Europe, and the Atlantic Islands. In like manner, eleven species of Bembidium form a group by themselves; and the Heteromera form two groups, one consisting of three genera and species of Opatridæ allied to a type found in Madeira, the other, Anthicodes, altogether peculiar.

Now each of these types may well be descended from a single species which originally reached the island from some other land; and the great variety of generic and specific forms into which some of them have diverged is an indication, and to some extent a measure, of the remoteness of their origin. The rich insect fauna of Miocene age found in Switzerland consists mostly of genera which still inhabit Europe, with others which now inhabit the Cape of Good Hope or the tropics of Africa and South America; and it is not at all improbable that the origin of the St. Helena fauna dates back to at least as remote, and not improbably to a still earlier, epoch. But if so, many difficulties in accounting for its origin will disappear. We know that at that time many of the animals and plants of the tropics, of North America, and even of Australia, inhabited Europe; while during the changes of climate, which, as we have seen, there is good reason to believe periodically occurred, there would be much migration from the temperate zones towards the equator, and the reverse. If, therefore, the nearest ally of any insular group now inhabits a particular country, we are not obliged to suppose that it reached the island from that country, since we know that most groups have ranged in past times over wider areas than they now inhabit. Neither are we limited to the means of transmission across the ocean that now exist, because we know that those means have varied greatly. During such extreme changes of conditions as are implied by glacial periods and by warm polar climates, great alterations of winds and of ocean-currents are inevitable, and these are, as we have already proved, the two great agencies by which the transmission of living things to oceanic islands has been brought about. At the present time the south-east trade-winds blow almost constantly at St. Helena, and the ocean-currents flow in the same direction, so that any transmission of insects by their means must almost certainly be from South Africa. Now there is undoubtedly a South African element in the insect-fauna, but there is no less clearly a European, or at least a north-temperate element, and this is very difficult to account for by causes now in action. But when we consider that this northern element is chiefly represented by remote generic affinity, and has therefore all the signs of great antiquity, we find a possible means of accounting for it. We have seen that during early Tertiary times an almost tropical climate extended far into the northern hemisphere, and a temperate climate to the Arctic regions. But if at this time (as is not improbable) the Antarctic regions were as much ice-clad as they are now it is certain that an enormous change must have been produced in the winds. Instead of a great difference of temperature between each pole and the equator, the difference would be mainly between one hemisphere and the other, and this might so disturb the trade winds as to bring St. Helena within the south temperate region of storms—a position corresponding to that of the Azores and Madeira in the North Atlantic, and thus subject it to violent gales from all points of the compass. At this remote epoch the mountains of equatorial Africa may have been more extensive than they are now, and may have served as intermediate stations by which some northern insects may have migrated to the southern hemisphere.

We must remember also that these peculiar forms are said to be northern only because their nearest allies are now found in the North Atlantic islands and Southern Europe; but it is not at all improbable that they are really widespread Miocene types, which have been preserved mainly in favourable insular stations. They may therefore have originally reached St. Helena from Southern Africa, or from some of the Atlantic islands, and may have been conveyed by oceanic currents as well as by winds. This is the more probable, as a large proportion of the St. Helena beetles live even in the perfect state within the stems of plants or trunks of trees, while the eggs and larvæ of a still larger number are likely to inhabit similar stations. Drift-wood might therefore be one of the most important agencies by which these insects reached the island.

Let us now see how far the distribution of other groups support the conclusions derived from a consideration of the beetles. The Hemiptera have been studied by Dr. F. Buchanan White, and though far less known than the beetles, indicate somewhat similar relations. Eight out of twenty-one genera are peculiar, and the thirteen other genera are for the most part widely distributed, while one of the peculiar genera is of African type. The other orders of insects have not been collected or studied with sufficient care to make it worth while to refer to them in detail; but the land-shells have been carefully collected and minutely described by Mr. Wollaston himself, and it is interesting to see how far they agree with the insects in their peculiarities and affinities.

Land-shells of St. Helena.—The total number of species is only twenty-nine, of which seven are common in Europe or the other Atlantic islands, and are no doubt recent introductions. Two others, though described as distinct, are so closely allied to European forms, that Mr. Wollaston thinks they have probably been introduced and have become slightly modified by new conditions of life; so that there remain exactly twenty species which may be considered truly indigenous. No less than thirteen of these, however, appear to be extinct, being now only found on the surface of the ground or in the surface soil in places where the native forests have been destroyed and the land not cultivated. These twenty peculiar species belong to the following genera: Hyalina (3 sp.), Patula (4 sp.), Bulimus (7 sp.), Subulina (3 sp.), Succinea (3 sp.); of which, one species of Hyalina, three of Patula, all the Bulimi, and two of Subulina are extinct. The three Hyalinas are allied to European species, but all the rest appear to be highly peculiar, and to have no near allies with the species of any other country. Two of the Bulimi (B. auris vulpinæ and B. darwinianus) are said to somewhat resemble Brazilian, New Zealand, and Solomon Island forms, while neither Bulimus nor Succinea occur at all in the Madeira group.

Omitting the species that have probably been introduced by human agency, we have here indications of a somewhat recent immigration of European types which may perhaps be referred to the glacial period; and a much more ancient immigration from unknown lands, which must certainly date back to Miocene, if not to Eocene, times.

Absence of Fresh-water Organisms.—A singular phenomenon is the total absence of indigenous aquatic forms of life in St. Helena. Not a single water-beetle or fresh-water shell has been discovered; neither do there seem to be any water-plants in the streams, except the common water-cress, one or two species of Cyperus, and the Australian Isapis prolifera. The same absence of fresh-water shells characterises the Azores, where, however, there is one indigenous water-beetle. In the Sandwich Islands also recent observations refer to the absence of water-beetles, though here there are a few fresh-water shells. It would appear therefore that the wide distribution of the same generic and specific forms which so generally characterises fresh-water organisms, and which has been so well illustrated by Mr. Darwin, has its limits in the very remote oceanic islands, owing to causes of which we are at present ignorant.

The other classes of animals in St. Helena need occupy us little. There are no indigenous mammals, reptiles, fresh-water fishes or true land-birds; but there is one species of wader—a small plover (Ægialitis sanctæ-helenæ)—very closely allied to a species found in South Africa, but presenting certain differences which entitle it to the rank of a peculiar species. The plants, however, are of especial interest from a geographical point of view, and we must devote a few pages to their consideration as supplementing the scanty materials afforded by the animal life, thus enabling us better to understand the biological relations and probable history of the island.

Native Vegetation of St. Helena.—Plants have certainly more varied and more effectual means of passing over wide tracts of ocean than any kinds of animals. Their seeds are often so minute, of such small specific gravity, or so furnished with downy or winged appendages, as to be carried by the wind for enormous distances. The bristles or hooked spines of many small fruits cause them to become easily attached to the feathers of aquatic birds, and they may thus be conveyed for thousands of miles by these pre-eminent wanderers; while many seeds are so protected by hard outer coats and dense inner albumen, that months of exposure to salt water does not prevent them from germinating, as proved by the West Indian seeds that reach the Azores or even the west coast of Scotland, and, what is more to the point, by the fact stated by Mr. Melliss, that large seeds which have floated from Madagascar or Mauritius round the Cape of Good Hope, have been thrown on the shores of St. Helena and have then sometimes germinated!

We have therefore little difficulty in understanding how the island was first stocked with vegetable forms. When it was so stocked (generally speaking), is equally clear. For as the peculiar coleopterous fauna, of which an important fragment remains, is mainly composed of species which are specially attached to certain groups of plants, we may be sure that the plants were there long before the insects could establish themselves. However ancient then is the insect fauna the flora must be more ancient still. It must also be remembered that plants, when once established in a suitable climate and soil, soon take possession of a country and occupy it almost to the complete exclusion of later immigrants. The fact of so many European weeds having overrun New Zealand and temperate North America may seem opposed to this statement, but it really is not so. For in both these cases the native vegetation has first been artificially removed by man and the ground cultivated; and there is no reason to believe that any similar effect would be produced by the scattering of any amount of foreign seed on ground already completely clothed with an indigenous vegetation. We might therefore conclude à priori, that the flora of such an island as St. Helena would be of an excessively ancient type, preserving for us in a slightly modified form examples of the vegetation of the globe at the time when the island first rose above the ocean. Let us see then what botanists tell us of its character and affinities.

The truly indigenous flowering plants are about fifty in number, besides twenty-six ferns. Forty of the former and ten of the latter are absolutely peculiar to the island, and, as Sir Joseph Hooker tells us, "with scarcely an exception, cannot be regarded as very close specific allies of any other plants at all. Seventeen of them belong to peculiar genera, and of the others, all differ so markedly as species from their congeners, that not one comes under the category of being an insular form of a continental species." The affinities of this flora are, Sir Joseph Hooker thinks, mainly African and especially South African, as indicated by the presence of the genera Phylica, Pelargonium, Mesembryanthemum, Oteospermum, and Wahlenbergia, which are eminently characteristic of southern extra-tropical Africa. The sixteen ferns which are not peculiar are common either to Africa, India, or America, a wide range sufficiently explained by the dust-like spores of ferns, capable of being carried to unknown distances by the wind, and the great stability of their generic and specific forms, many of those found in the Miocene deposits of Switzerland, being hardly distinguishable from living species. This shows, that identity of species of ferns between St. Helena and distant countries does not necessarily imply a recent origin.

The Relation of the St. Helena Compositæ.—In an elaborate paper on the Compositæ, Mr. Bentham gives us some valuable remarks on the affinities of the seven endemic species belonging to the genera Commidendron, Melanodendron, Petrobium, and Pisiadia, which forms so important a portion of the existing flora of St. Helena. He says: "Although nearer to Africa than to any other continent, those composite denizens which bear evidence of the greatest antiquity have their affinities for the most part in South America, while the colonists of a more recent character are South African." ... "Commidendron and Melanodendron are among the woody Asteroid forms exemplified in the Andine Diplostephium, and in the Australian Olearia. Petrobium is one of three genera, remains of a group probably of great antiquity, of which the two others are Podanthus in Chile and Astemma in the Andes. The Pisiadia is an endemic species of a genus otherwise Mascarene or of Eastern Africa, presenting a geographical connection analogous to that of the St. Helena Melhaniæ, with the Mascarene Trochetia."

Whenever such remote and singular cases of geographical affinity as the above are pointed out, the first impression is to imagine some mode by which a communication between the distant countries implicated might be effected; and this way of viewing the problem is almost universally adopted, even by naturalists. But if the principles laid down in this work and in my Geographical Distribution of Animals are sound, such a course is very unphilosophical. For, on the theory of evolution, nothing can be more certain than that groups now broken up and detached were once continuous, and that fragmentary groups and isolated forms are but the relics of once widespread types, which have been preserved in a few localities where the physical conditions were especially favourable, or where organic competition was less severe. The true explanation of all such remote geographical affinities is, that they date back to a time when the ancestral group of which they are the common descendants had a wider or a different distribution; and they no more imply any closer connection between the distant countries the allied forms now inhabit, than does the existence of living Equidæ in South Africa and extinct Equidæ in the Pliocene deposits of the Pampas, imply a continent bridging the South Atlantic to allow of their easy communication.

Concluding Remarks on St. Helena.—The sketch we have now given of the chief members of the indigenous fauna and flora of St. Helena shows, that by means of the knowledge we have obtained of past changes in the physical history of the earth, and of the various modes by which organisms are conveyed across the ocean, all the more important facts become readily intelligible. We have here an island of small size and great antiquity, very distant from every other land, and probably at no time very much less distant from surrounding continents, which became stocked by chance immigrants from other countries at some remote epoch, and which has preserved many of their more or less modified descendants to the present time. When first visited by civilised man it was in all probability far more richly stocked with plants and animals, forming a kind of natural museum or vivarium in which ancient types, perhaps dating back to the Miocene period, or even earlier, had been saved from the destruction which has overtaken their allies on the great continents. Unfortunately many, we do not know how many, of these forms have been exterminated by the carelessness and improvidence of its civilised but ignorant rulers; and it is only by the extreme ruggedness and inaccessibility of its peaks and crater-ridges that the scanty fragments have escaped by which alone we are able to obtain a glimpse of this interesting chapter in the life-history of our earth.

119 St. Helena: a Physical, Historical, and Topographical Description of the Island, &c. By John Charles Melliss, F.G.S., &c. London: 1875.

120 Mr. Marsh in his interesting work entitled The Earth as Modified by Human Action (p. 51), thus remarks on the effect of browsing quadrupeds in destroying and checking woody vegetation.—"I am convinced that forests would soon cover many parts of the Arabian and African deserts if man and domestic animals, especially the goat and the camel, were banished from them. The hard palate and tongue, and strong teeth and jaws of this latter quadruped enable him to break off and masticate tough and thorny branches as large as the finger. He is particularly fond of the smaller twigs, leaves, and seed-pods of the Sont and other acacias, which, like the American robinia, thrive well on dry and sandy soils, and he spares no tree the branches of which are within his reach, except, if I remember right, the tamarisk that produces manna. Young trees sprout plentifully around the springs and along the winter water-courses of the desert, and these are just the halting stations of the caravans and their routes of travel. In the shade of these trees annual grasses and perennial shrubs shoot up, but are mown down by the hungry cattle of the Bedouin as fast as they grow. A few years of undisturbed vegetation would suffice to cover such points with groves, and these would gradually extend themselves over soils where now scarcely any green thing but the bitter colocynth and the poisonous foxglove is ever seen."

121 Coleoptera Sanctæ Helenæ, 1877; Testacea Atlantica, 1878.

122 On Petermann's map of Africa, in Stieler's Hand-Atlas (1879), the Island of Ascension is shown as seated on a much larger and shallower submarine bank than St. Helena. The 1,000 fathom line round Ascension encloses an oval space 170 miles long by 70 wide, and even the 300 fathom line, one over 60 miles long; and it is therefore probable that a much larger island once occupied this site. Now Ascension is nearly equidistant between St. Helena and Liberia, and such an island might have served as an intermediate station through which many of the immigrants to St. Helena passed. As the distances are hardly greater than in the case of the Azores, this removes whatever difficulty may have been felt of the possibility of any organisms reaching so remote an island. The present island of Ascension is probably only the summit of a huge volcanic mass, and any remnant of the original fauna and flora it might have preserved may have been destroyed by great volcanic eruptions. Mr. Darwin collected some masses of tufa which were found to be mainly organic, containing, besides remains of fresh-water infusoria, the siliceous tissue of plants! In the light of the great extent of the submarine bank on which the island stands, Mr. Darwin's remark, that—"we may feel sure, that at some former epoch, the climate and productions of Ascension were very different from what they are now,"—has received a striking confirmation. (See Naturalist's Voyage Round the World, p. 495.)

123 "Notes on the Classification, History, and Geographical Distribution of Compositæ."—Journal of the Linnean Society, Vol. XIII. p. 563 (1873).

124 The Melhaniæ comprise the two finest timber trees of St. Helena, now almost extinct, the redwood and native ebony.

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 Rh the eruption of Vesuvius"; the following from Æschylus: "Raising the Ghost of Darius" ; "Atossa's Dream " ; "The Furies." I hope they exist in a perfect state; and if they do, they are all well worth etching in a bold manner, which I think Blake is likely to do with great success, and perhaps at an expense that will not be burthensome. But, at any rate, give him one to do first for a trial. The exhibition of a painter s noblest sentiments and grandest thoughts must certainly become as striking and interesting in his life as their several poems in the lives of Milton, Homer, or Virgil. I am glad you are satisfied with the introduction; in this you have had the success of a friend both zealous and skilful. I confess, great as my regard was for the man, to write his life and speak the truth without offence seems attended with considerable difficulty. I am not at all surprised that many passages in this eminent man's letters were truly eloquent and beautiful. Indeed I should have been confounded had they not been so; because whatever advantages


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 136 can bestow, they are but so many modifications of the light of the understanding and the feelings and affections of the heart. I am sure Romney's memory will want no grace or decoration which your pen cannot give, and therefore any effort of mine would be comparatively poor and tedious, like "the shuffling gait of a tired nag." Yet I am not so churlish that I would not lend an endeavour, however weak, to honour a departed friend; but even this must be independent, and not comparative. I do not remember Mr. Robinson, but I rejoice in his success and the inspiration of his son. The works I am employed on at present are Earl Howe and Captain Montague. I have troubled you, by Mr. Blake, with a short tract written for Dr. Rees's Cyclopedia, on "Basso Relievo," with one of the prints referred to at the end of the article; the rest are not yet engraven.

I do not send it from the vanity of giving information to you, because I daresay you are well acquainted with all it contains: but because I would not publish anything of this kind without sending a copy, even though it should not be worth your notice. A happy release from his afflictions to poor Blake; and to you, my dear friend, many happy years unclouded by misfortune


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 pr. n. m. Abnimos, Nimos, a gentile philosopher, friend of R. Meïr [prob. identical with the cynic philosopher Oenomaus of Gadara]. Gen. R. s. 65; a. e.. Ḥag. 15$b$.

Targ. Y. II Deut. XIV, 18, read with Y. I, v..

f. understanding, speculation. Meg. 24$b$ (Ms. M. ) it depends on the speculative faculty (not on the physical sight). Ab. Zar. 28$b$ (Ms. M., cmp. Tosaf. a. l.) an affection of the eye-sight is connected with (has influence on) the mental faculties; (oth. opin., cmp. , the fat surrounding the heart).

(b. h.; √, cmp. ) to stuff; to fatten, feed (act. a. neut.) B. Mets 86$b$ (expl. ăbusim, I Kings V, 3) ) which people fatten with force. Ib. that stand feeding as they please. Sabb. XXIV, 3 (155$b$)  you must not (on the Sabbath) stuff the camel; expl. ib. you must not make  a manger of her stomach (fill up to swelling); a. fr.—Part. pass.  (=). Meg. 9$a$, a. e. (one of the changes said to have been made by the authors of the Septuag.).

unbewitched! may no harm befall you! Y. Ab. Zar. I, end, 40$b$ he did not say abascanta, but etc. Y. Ber. IV, 13$c$ top (corr. acc.).

v..

m. pl. blains, pustules. Targ. O. Ex. IX, 9 (Var. f. pl.).

, v..

f. search, begging, the appearance of the poor for their share in the crop. Pl. . Peah IV, 5 (Y. ed. IV, 3 ) three times a day the poor would come (cmp. etym. Y. ib. 18$b$ top). [Oth. comment. ref. to etym. in Y. l. c. a. Targ. Obad. v. 6: "the owner appears" &c.]

m. (=, cmp. ) tin. Targ. O. Num. XXXI, 22.

m. (b. h.; v., cmp. ), (thick, whirling) dust, powder. Sabb. III, 3 the (heated) sand on the roads. Ḥull. 91$a$. Cant. R. to III, 6; a. fr.— the refuse of writing material, or the colored sand strewn over the writing. Sabb. XII, 5 if one writes (on the Sabb.) with a fluid or sap of fruits (instead of ink), or in the sand on roads or in the writer's powder.—Trnsf. (cmp. ) connection, something akin to, shade of, as a shade of slander;  a shade of usury;  an agricultural occupation indirectly related to those forbidden in the Sabbath year; v. infr.-Pl. . Tosef. Ab. Zar. I, 10 in (ed. Zuck. ) the word abak in its figur. sense is applied to four things; cmp. B. Bath. 165$a$; B. Mets, 61$b$; 67$a$; Succ. 40$b$.

Pi. , (denom. of foreg.) to cover with powder, esp. plants, for fertilizing.. Shebi. II, 2 (cmp. Y. Gem. a. l.). M. Kat. 3$a$. Y. Sabb. VII, 10$a$ top.—Part. pass., powdered. Y. Bicc. I, 63$d$ bot. (read ) grapes fertilized with powder. [Ar. "to remove the dust"(?)]

Hithpa. a. Nithpa. ,, to be covered, or cover one's self with dust. Gen. R. s. 43.—Metaph. to sit at one's feet as a disciple. Aboth. I, 4.

(√,, cmp. , ) 1) to entangle, twist, twine. Men. 42$a$ (perh. Pa.) he twined (the show-fringes) with loops.—2) (neut. v.) to be attached to, cling to (idolatry etc.). Snh. 64$a$; Ab. Zar. 14$b$. Ib. 17$a$ he was very deeply attached to sensuality.

ch.=h. . Targ. Ex. IX, 9; a. e.

Tosef. Mikv. V, 7, read.

v. ch.

v. h.

f. (b. h.; v. ) spices, spice-box. Fig. (peddlar's spice-box) a great scholar. Cant. R. to III, 6 end.

pr. n. m. Eucolus, father of R. Zechariah. Git. 56$a$; Lam. R. to IV, 2. Tosef. Sabb. XVI (XVII), 6 (Var., ). Cmp. .

Men. 33$a$, read , v. next w.

f. pl. loops, leather rings, on bedsteads for the reception of cords; in door cases, for hanging doors in. Ned. 56$b$; Snh. 20$b$. a couch is called dargesh, when it is carried in and out (to be put up and taken apart) by means of loops (through which the cords are fastened); opp. mittah, v. .—Men. 33$a$, Erub. 11$b$ ed. (Ms. M., v. Rabb. D. S. a. l. note), explain. 'an indication of hinges'.—Macc. 23$a$ (loops in the punishing scourge).

or Hif. , v..

(√, v. ; cmp., ) to be bent, pressed, thick.

Pi. , 1) to strengthen, harden (cmp. ). Snh. 109$b$ (play on Abiram, Num. XVI, 1) Ms. M. (Rashi, ed. ) he hardened his heart against repentance.— 2) (denom. of ) to measure wings, to define city limits, for Sabbath distances, in cases of wing-like projections beyond the line. Erub. V, 1 (accord. to Rab's spelling, while Sam. read, v. Y. ib. 22$b$, Bab. ib. 53$a$) how do we measure outskirts of a city in order to draw the Sabbath line?; v. etymol. definit. Y. a. Babl. 11. cc. a. Y. Ber. VII, 12$c$ top.—3) (b. h. Hif.) to soar, take wings. Gen. R. s. 42 (play on Shemeber, Gen. XIV, 2) Ar. s. v.  (ed. ) he took wings to fly and obtain wealth.


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Rh much of this irresponsibility and detachment to the last. It is true that after much search he found, or believed that he found, certain general laws, or principles, to which his observations could be attached, under which they could be grouped. But one often feels, in reading his opening chapters, that they are a sham façade, giving a deceptive appearance of unity to a complicated and irregular set of buildings, richly stored with miscellaneous objects of interest. His doctrine of the relativity of laws, which is the foundation of enlightened conservatism, and has been used in defence of much conservatism which is not enlightened, is not a sufficient foundation for a constructive system, but was an admirable starting-point for a man whose primary interest lay in observing and comparing different institutions and drawing inferences from their similarities and diversities. 'Any one who has eyes to see,' he wrote in his subsequent Defence of the Spirit of Laws, 'must see at a glance that the object of the work was the different laws, customs and usages of the peoples of the world.' A vast, an overwhelming subject, which the author failed to succeed in mastering and controlling, or bringing within a synthetic grasp. And owing to this failure the Spirit of Laws has been not unfairly described as being, not a great book, but the fragments of a great book. What he did succeed in

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28 doing was in indicating the path by which alone effective and fruitful progress could be made either in jurisprudence or in the science of politics, the path through diversity to uniformity, through facts to principles. He refashioned political science and made it a science of observation, and by so doing he made the same new departure in political and legal science as Bacon had made before him in physical science. He closed the period of the schoolmen. He was not content to mumble the dry bones of Roman law. He turned men away from abstract and barren speculations to the study and comparison of concrete institutions. And it is in this sense that he may be claimed as one of

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